Psychomyioidea

Ecnomina sp., from here.

Belongs within: Trichoptera.
Contains: Polycentropodidae.

The Psychomyioidea are a clade of caddisflies characterised by flattened larvae with a hydrophobous cuticle (Ivanov & Sukatsheva 2002).

Synapomorphies (from Ivanova & Sukatsheva 2002): Larval pronotal hind angles joining sternum behind coxa, prelabio-hypopharynx sclerotised dorsally, prementum not distinguishable, spinneret tending to be long; larva flat, larval cuticle hydrophobous; antenna with one basiconic sensilla; adult scutal warts rounded.

<==Psychomyioidea
    |--Dysoneuridae [Disoneuridae] GE05
    |    |--Onkovena sharategensis IS02
    |    |--Bullvenia grandis IS02
    |    `--Utania remissa Sukacheva 1990 RJ93
    `--+--+--Xiphocentronidae GE05
       |  `--Psychomyiinae [Psychomyidae, Psychomyiidae] IS02
       |       |--Psychomyia Latreille 1829 BZ-M04
       |       |    |--P. flavida KP19
       |       |    `--P. pusilla (Fabricius 1781) BZ-M04
       |       |--Lype McLachlan 1878 BZ-M04
       |       |    |--L. reducta (Hagen 1868) BZ-M04
       |       |    `--L. sericea RJ93
       |       |--Metalype Klapálek 1898 BZ-M04
       |       |    `--M. fragilis (Pictet 1834) BZ-M04
       |       `--Tinodes Curtis 1834 BZ-M04
       |            |--T. algiricus McLachlan 1880 BZ-M04
       |            |--T. assimilis McLachlan 1865 BZ-M04
       |            |--T. baenai González & Otero 1984 BZ-M04
       |            |--T. dives (Pictet 1834) BZ-M04
       |            |--T. maclachlani Kimmins 1966 BZ-M04
       |            |--T. maculicornis (Pictet 1834) BZ-M04
       |            |--T. maroccanus Mosely 1938 BZ-M04
       |            `--T. waeneri (Linnaeus 1758) BZ-M04
       `--+--Dipseudopsidae GE05
          `--+--Electrabertidae GE05
             |--Polycentropodidae GE05
             `--Ecnomidae GE05
                  |--Archaeotinodes angusta RJ93
                  |--Ecnomus McLachlan 1864 BZ-M04
                  |    |--*E. tenellus (Rambur 1842) [=Philopotamus tenellus] N83
                  |    |--E. continentalis Ulmer 1916 N83
                  |    |--E. cygnitus Neboiss 1982 N83
                  |    |--E. deceptor McLachlan 1884 BZ-M04
                  |    |--E. pansus Neboiss 1982 N83
                  |    |--E. russellius Neboiss 1977 N83
                  |    |--E. tillyardi Mosely 1953 N83
                  |    `--E. turgidus Neboiss 1982 N83
                  `--Ecnomina Kimmins 1953 N83
                       |--*E. spinosa Kimmins 1953 N83
                       |--E. batyle Neboiss 1977 N83
                       |--E. chorisis Neboiss 1978 N83
                       |--E. cohibilis Neboiss 1982 N83
                       |--E. irrorata Kimmins 1953 N83
                       |--E. krokale Neboiss 1978 N83
                       |--E. legula Neboiss 1977 N83
                       |--E. merga Neboiss 1982 N83
                       |--E. mesembria Neboiss 1982 N83
                       |--E. scindens Neboiss 1982 N83
                       |--E. sentosa Neboiss 1982 N83
                       |--E. thinotes Neboiss 1978 N83
                       |--E. trifurcata Kimmins 1953 N83
                       |--E. trulla Neboiss 1982 N83
                       |--E. vega Neboiss 1977 N83
                       `--E. viatica Neboiss 1982 N83

*Type species of generic name indicated

REFERENCES

[BZ-M04] Bonada, N., C. Zamora-Muñoz, M. Rieradevall & N. Prat. 2004. Trichoptera (Insecta) collected in Mediterranean river basins of the Iberian Peninsula: taxonomic remarks and notes on ecology. Graellsia 60 (1): 41–69.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[IS02] Ivanov, V. D., & I. D. Sukatsheva. 2002. Order Trichoptera Kirby, 1813. The caddisflies (=Phryganeida Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 199–220. Kluwer Academic Publishers: Dordrecht.

[KP19] Kawahara, A. Y., D. Plotkin, M. Espeland, K. Meusemann, E. F. A. Toussaint, A. Donath, F. Gimnich, P. B. Frandsen, A. Zwick, M. dos Reis, J. R. Barber, R. S. Peters, S. Liu, X. Zhou, C. Mayer, L. Podsiadlowski, C. Storer, J. E. Yack, B. Misof & J. W. Breinholt. 2019. Phylogenomics reveals the evolutionary timing and pattern of butterflies and moths. Proceedings of the National Academy of Sciences of the USA 116 (45): 22657–22663.

[N83] Neboiss, A. 1983. Checklist and bibliography of the Australian caddis-flies (Trichoptera). Australian Society for Limnology Special Publication 5: 1–132.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.