Belongs within: Phalangiidae.
Zacheus is a genus of large-bodied phalangiid harvestmen found in the eastern Mediterranean and western Asia (Kurt et al. 2015). Historically, there has been confusion regarding the correct spelling of the genus name; Crawford (1992) provides reasoning for use of the spelling 'Zachaeus'.
Characters (from Kurt et al. 2015): Body large, heavily denticulated dorsally; chelicerae usually strong, second segment enlarged; pedipalps normally structured, strong, robust; legs short, first pair much thicker than others; truncus of penis basally widened, parallel-sided on distal half, distally shallow spoon-shaped, glans usually banana-shaped (shorter in Z. anatolicus and Z. seyyari), stylus long.
<==Zachaeus Koch 1839 K14 [=Zacheus Koch 1839 C92]
|--Z. crista (Brullé 1832) K14 (see below for synonymy)
|--Z. anatolicus (Kulczyński 1903) K14 [=Egaenus crista var. anatolicus R23, Z. crista var. anatolicus R23]
|--Z. birulai Redikorzev 1936 S99
|--Z. canaliculatus Kollar in Roewer 1911 [=Phalangium canaliculatum ms] R23
|--Z. caucasicus Roewer 1911 R23
|--Z. cubanus Roewer 1923 R23
|--‘Dasylobus’ egaenoides Simon 1885 (n. d.) R23
|--Z. hebraicus (Simon 1884) K14 [=Phalangium hebraicum R23, Z. crista var. hebraicus R23]
|--Z. kervillei (Sørensen 1912) (n. d.) KKY15 [=Egaenus kervillei R23]
|--Z. leucomelas (Simon 1884) R23 [=Phalangium hebraicum leucomelas R11, Zachaeus crista var. leucomelas R23]
|--Z. lupatus (Eichwald 1830) [incl. E. gulosus Simon 1878] S03
|--Z. macrinus Roewer 1956 R56
|--Z. mirabilis (Caporiacco 1949) KKY15
|--Z. orchimonti (Giltay 1933) K14
|--Z. palpipes Roewer 1915 R23
|--‘Opilio’ punctipes Koch 1878 (n. d.) R23
|--Z. redikorzevi (Staręga & Chevrizov 1978) K14
|--Z. seyyari Kurt, Koç & Yağmur 2015 KKY15
|--Z. shachdag (Snegovaya & Staręga 2008) KKY15
`--Z. simferopolensis Chemeris & Kovblyuk 2005 [=Z. simpheropolensis] CK05
Zachaeus crista (Brullé 1832) K14 [=Phalangium crista C92, Egaenus crista C92; incl. E. hungaricus Lendl 1894 M78, Zachaeus crista var. hungaricus R23, Paropilio lineatus Roewer 1956 M78, *Zachaeus mordax Koch 1839 C92, Egaenus mordax R23, Z. trinotatus Koch 1839 R23, E. variegatus Lendl 1894 M78, Zachaeus crista var. variegatus R23]
*Type species of generic name indicated
REFERENCES
[CK05] Chemeris, A. N., & N. M. Kovblyuk. 2005. A contribution to the knowledge of the harvestman fauna of the Crimea (Arachnida: Opiliones). Arthropoda Selecta 14 (4): 305–328.
[C92] Crawford, R. L. 1992. Catalogue of the genera and type species of the harvestman superfamily Phalangioidea (Arachnida). Burke Museum Contributions in Anthropology and Natural History 8: 1–60.
[K14] Kurt, K. 2014. Updated checklist of harvestmen (Arachnida: Opiliones) in Turkey. Arch. Biol. Sci. 66 (4): 1617–1631.
[KKY15] Kurt, K., H. Koç & E. A. Yağmur. 2015. A new species of Zachaeus C. L. Koch from Turkey (Opiliones, Phalangiidae). ZooKeys 514: 15–23.
[M78] Martens, J. 1978. Spinnentiere, Arachnida: Weberknechte, Opiliones. Gustav Fischer Verlag: Jena.
[R11] Roewer, C.-F. 1911. Übersicht der Genera der Subfamilie der Phalangiini der Opiliones Palpatores nebst Beschreibung einiger neuer Gattungen und Arten. Archiv für Naturgesichte 77 (Suppl. 2): 1–106.
[R23] Roewer, C.-F. 1923. Die Weberknechte der Erde: Systematische Bearbeitung der bisher bekannnten Opiliones. Gustav Fischer: Jena.
[R56] Roewer, C. F. 1956. Über Phalangiinae (Phalangiidae, Opiliones Palpatores). (Weitere Weberknechte XIX). Senckenbergiana Biologica 37 (3–4): 247–318.
[S99] Snegovaya, N. 1999. Contribution to the harvest spider (Arachnida, Opiliones) fauna of the Caucasus. Tr. J. of Zoology 23: 453–459.
[S03] Staręga, W. 2003. On the identity and synonymies of some Asiatic Opilioninae (Opiliones: Phalangiidae). Acta Arachnologica 52 (2): 91–102.
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