Belongs within: Mollusca.
Contains: Scoliostomatidae, Caseolus, Leiostyla, Clisospiroidea, Bellerophontoidea, Archinacelloidea, Pelagielloidea, Macluritida, Neomphaliones, Patellogastropoda, Vetigastropoda, Neritimorpha, Loxonematoidea, Heterobranchia, Peruneloidea, Cyclophoroidea, Ampullariidae, Viviparoidea, Orthonematoidea, Subulitoidea, Soleniscidae, Meekospiridae, Cerithiimorpha, Hypsogastropoda.
The Gastropoda, snails and slugs, are a major clade of molluscs characterised by the process of torsion during larval development with the visceral mass and mantle becoming turned 180° relative to the head and foot. The shell (if present) has a single valve and is often spirally coiled; an operculum is also often present that closes the aperture of the shell when the body is withdrawn. Both of these features may be modified in derived subgroups: the shell may be lost or coiling reduced, and torsion may be secondarily reversed over the course of continued development. Gastropods are ancestrally aquatic but a number of derived lineages have become terrestrial. Embryonic development in aquatic gastropods may be indirect, with a free-living, planktotrophic larval stage, or direct, with lecithotrophic embryos hatching into crawling juveniles. These differing modes of development are often reflected in different aspects of the protoconch (the initial, larval section of the shell which often differs in appearance from the mature teleoconch) with planktotrophic developers have more whorls to the protoconch than lecithotrophs.
Historically, gastropods have often been divided between three major groupings—the Prosobranchia (possessing a gill and torted anatomy at maturity), Opisthobranchia (gilled but with torsion not evident at maturity), and Pulmonata (respiring via a lung)—but the 'prosobranchs' are recognised as paraphyletic to the last two groups (which together form part of the broader clade Heterobranchia). The 'prosobranchs' have in turn been historically divided between the grades Archaeogastropoda, Mesogastropoda and Neogastropoda but, again, the 'archaeogastropods' are now regarded as paraphyletic with the Neogastropoda and most 'mesogastropods' forming the clade Caenogastropoda. The Caenogastropoda and Heterobranchia together form a clade Apogastropoda, supported by a number of synapomorphies such as the presence of a single columellar muscle, loss of the right auricle, and the oesophagus originating at the posterior end of the buccal mass (Simone 2011).
Various Early Palaeozoic fossils such as Bellerophontoidea and Pelagielloidea resemble modern gastropods in the possession of a coiled shell but have not been universally accepted as gastropods due to uncertainty whether or not they exhibited torsion. Thomas et al. (2020) argued for the presence of torsion in Pelagielloidea on the basis of functional interpretations of preserved chaetae and internal grooves on the shell.
<==Gastropoda (see below for synonymy)
|--+--Pelagielloidea TRM20
| `--Macluritida TRM20
`--+--Neomphaliones BR17
`--+--+--Patellogastropoda BR17
| `--Vetigastropoda S11
`--Adenogonogastropoda [Branchiata, Ectophthalma, Teleohydrophila] S11
|--Neritimorpha S11
`--Apogastropoda (see below for synonymy) S11
| i. s.: Protostylus Mansuy 1914 (n. d.) KC60
| `--*P. lantenoisi Mansuy 1914 KC60
| ‘Cyclora’ imbricata Whiteaves 1881 G88
| ‘Cyclora’ valvatiformis Whiteaves 1881 G88
|--Loxonematoidea PC08
`--+--Heterobranchia S11
`--Caenogastropoda (see below for synonymy) S11
| i. s.: Gaeatrivia millardi Cate 1979 M03
| Serratovolva dondani Cate 1964 M03
| Decorochilina Frýda 1999 F01
| Plicatus Pan & Erwin 2002 [Plicatusidae] BR05
| |--*P. scalaris Pan & Erwin 2002 PE02
| `--P. multifilaris Pan & Erwin 2002 PE02
| Spanionema Whidborne 1891 [Spanionematidae, Spanionematoidea] BR05
| `--*S. scalaroides (Whidborne 1889) [=Loxonema scalaroides] KC60
| Spirochrysalis Kittl 1894 NE02
| `--S. protomicra Pan 1982 NE02
| Lampanella minima PC08
| Pragoscutula Frýda 1998 PC08, BR05 [Pragoscutulidae]
| `--P. wareni Frýda 1998 FB01
| Dendropupoidea [Anthracopupoidea, Dendropupinae, Procyclophoroida] BR05
| |--Maturipupa Pilsbry 1926 KC60
| | `--*M. vermilionensis (Bradley 1872) [=Pupa vermilionensis] KC60
| |--Anthracopupa Whitfield 1881 [Anthracopupidae, Anthracopupinae] BR05
| | |--*A. ohioensis Whitfield 1881 KC60
| | `--A. britannica Cox 1926 TTE93
| `--Dendropupa Owen 1859 BR17 (see below for synonymy)
| |--*D. vetusta (Dawson 1859) [=Pupa vetusta] KC60
| |--D. grandaeva (Dawson 1880) TTE93
| `--D. primaeva (Matthew 1895) TTE93
| Acanthonema Grabau in Sherzer & Grabau 1909 BR05 (see below for synonymy)
| `--A. newberryi (Meek 1871) TTE93 (see below for synonymy)
| Ampezzanildidae BR17
| |--Cassianilda Bandel 1995 NE02
| |--Stuorilda Bandel 1995 NE02
| `--Ampezzanilda Bandel 1994 BR05
| `--*A. aialensis (Zardini 1980) [=Promathildia aialensis] BR17
| Kittlidiscus Haas 1953 BR05 (see below for synonymy)
| `--K. bronni (Klipstein 1845) KC60 (see below for synonymy)
| Spirostylidae [Spirostylinidae] TE01
| |--Spirostylus Kittl 1894 TE01
| | |--*S. subcolumnaris (Münster 1841) [=Melania subcolumnaris] TE01
| | `--S. linctus (Böhm 1895) (see below for synonymy) TE01
| `--Climacina TTE93
| |--C. catharinae Gemmellaro 1876 TTE93
| `--C. mariae Gemmellaro 1876 TTE93
| Peruneloidea BR17
|--Cyclophoroidea S11
`--Hydrogastropoda S11
|--Ampullariidae S11
`--Epiarthroidea [Teleobranchia, Vivipariformes, Vivipariformii] S11
|--Viviparoidea S11
`--Sorbeoconcha (see below for synonymy) S11
| i. s.: Globocornus Espinosa & Ortea 2010 [Globocornidae] BR17
| `--*G. darwini Espinosa & Ortea 2010 BR17
| Acteonina d’Orbigny 1850 BR05 (see below for synonymy)
| |--*A. carbonaria (de Koninck 1843) [=Chemnitzia carbonaria] BR05
| |--A. acuta (d’Orbigny 1841) [=Tornatina acuta] BR17
| `--A. obesa Stoliczka 1868 TTE93
| Orthonematoidea BR17
| Subulitina [Subulitata] GS75
| |--Subulitoidea GS75
| `--Soleniscoidea BR17
| |--Soleniscidae BR17
| |--Meekospiridae BR17
| `--Anozygidae BR17
| |--Anozyga Hoare 1980 [Anozyginae] BR05
| | `--*A. bulla Hoare 1980 BR17
| `--Tmetonema Longstaff 1912 [Tmetoneminae] BR05
| `--*T. subsulcatum Longstaff 1912 KC60
| Pluviostilla BR17
|--Cerithiimorpha S11
`--Hypsogastropoda S11
Gastropoda incertae sedis:
Spiniplatyceras FBL02
Serpentubina Horný 1964 FBL02
Kiviasukkaan Peel 1975 FBL02
Pragoserpulina Frýda 1998 [Pragoserpulinidae] BR05
`--*P. tomasi Frýda 1998 BR17
Luoguispira Pan & Erwin 2002 PE02
|--*L. micra Pan & Erwin 2002 PE02
|--L. lineata Pan & Erwin 2002 PE02
`--L. multilinea Pan & Erwin 2002 PE02
Luoguella Pan & Erwin 2002 PE02
|--*L. elegantula Pan & Erwin 2002 [=Gyrospira elegantula] PE02
`--L. diana Pan & Erwin 2002 [=Gyrospira diana] PE02
Propupa PE02
Extendilabrum PE02
Tetrabubispira PE02
Kalchreuthia Gründel & Nützel 1998 GK02
Paladilha (Lartella Cossmann 1921) GK02
Pseudotaphrus (Pezantia Cossmann 1896) GK02
Chlamydarion P02
Derjuginella rufofasciata GAS03
Erheria vincta GAS03
Melanochlamis GAS03
|--M. cylindrica HS01
`--M. diomedea GAS03
Yokoyamia ornatissima GAS03
Epheria turrita KBC03
Tersiella plicosa KBC03
Ausola angustata KBC03
Hinia cuvieri M62
Neritula donovani M62
Oxytrema Rafinesque 1819 B63
`--O. silicula B96
Hystrivasum horridum Heilprin 1886 HP04
Benthonella tenella (Jeffreys 1869) OT04
Chlupacispira spinosa Blodgett & Rohr 1989 FB04
Eulepetopsis vitrea GO06
Purkynespira Frýda & Bandel 1997 FB01
`--P. reticulata FB01
Streptopelma henchmani Marwick 1926 F26
Latirofusus Cossmann 1889 F27
|--*L. funiculosus F27
|--L. aciformis [=Fusus aciformis] F27
|--L. acus [=Fusus acus] F27
|--L. lancea [=Fusus lancea] F27
|--L. optatus M. & M. 1923 F26
`--L. spiceri (Tenison-Woods 1877) [=Fusus spiceri; incl. L. nigrofuscus Tate 1891] F27
Buccinulus F26
|--B. albus F26
|--B. huttoni Kirk 1882 F26
`--B. kirki Hutton 1873 [incl. B. gracilis Kirk 1882] F26
Triploca Tate 1894 F26
|--T. ligata F26
`--T. waihaoensis Marshall & Murdoch 1923 F26
Leucotina F26
|--L. casta (Adams 1853) [=Monoptgyma casta] H09
|--L. helva Hedley 1900 H09
`--L. pura (Adams 1855) F26
Sublacuna Cossmann 1899 [=Strebloramphus Tate 1898 (preoc.)] F27
`--*S. mirula (Tate 1898) [=*Strebloramphus mirulus] F27
Coptochetus asperulus [=Fusus asperulus] F27
Trifora Deshayes in Blainville 1828 O27b
|--*T. perversa O27b
|--T. callipyrga Bartsch 1907 O27b
|--T. carpenteri Bartsch 1903 O27b
|--T. catalinensis Bartsch 1907 O27b
|--T. montereyensis Bartsch 1907 O27b
|--T. pedroana Bartsch 1907 O27b
|--T. peninsularis Bartsch 1907 O27b
`--T. stearnsi Bartsch 1907 O27b
Barocospira Horný 1964 FBL02
Ibergia Blodgett & Frýda 1999 FBL02
Crassimarginatidae BR05
|--Yukonoconcha Frýda, Blodgett & Lenz 2002 FBL02
| `--*Y. pedderi Frýda, Blodgett & Lenz 2002 FBL02
`--Crassimarginata Jhaveri 1969 BR05
|--*C. crassicosta Jhaveri 1969 FBL02
`--C. tenuicosta Jhaveri 1969 FBL02
Scoliostomatidae BR05
Levihelix pusilla Gründel 2000 GK02
Sinuginella pygmaea HS01
Adelphotectonica reevi HS01
Bouchetriphora pallida HS01
Limulatys reliquus HS01
Pisinna HS01
|--P. manawatawhia HS01
|--P. olivacea HS01
| |--P. o. olivacea HS01
| `--P. o. impressa HS01
|--P. rekohuana HS01
| |--P. r. rekohuana HS01
| `--P. r. lactorubra HS01
|--P. semiplicata HS01
`--P. zosterophila HS01
Sagenotriphora ampulla HS01
Camita rotellina (Gould 1860) [=Trochus (Monodonata) rotellinus] H08
Turcia H09
|--T. maculata (Brazier 1877) [=Thalotia maculata] H09
|--T. monilifera Adams 1855 H09
`--T. montrouzieri (Fischer 1878) [=Tectaria montrouzieri] H09
Mecoliotia spinosa Hedley 1902 H09
Quoyia decollata (Quoy & Gaimard 1833) [=Planaxis decollatus] H09
Eglisia tricarinata Adams & Reeve 1840 H09
Couthouyia aculeata Hedley 1900 H09
Oscilla H09
|--O. tasmanica (Ten.-Woods 1877) [=Parthenia tasmanica] H09
`--O. terebellum [=Cingula terebellum; incl. O. exarata, Parthenia exarata] C64
Elusa subulata (Adams 1855) [=Pyramidella subulata] H09
Scalenostoma striatum Hedley 1900 H09
Distortrix decipiens (Reeve 1844) [=Triton decipiens] H09
Laimodonta conica Pease 1862 H09
Paraplysia piperata (Smith 1884) [=Aplysia piperata] H09
Euselenops luniceps (Cuvier 1830) [=Pleurobranchus luniceps] H09
Placomopherus insignis Smith 1884 H09
Adiozoptyxis Dietrich 1925 [Adiozoptyxinae, Adiozoptyxisinae] BR05
`--*A. polymorpha (Gemmellaro 1865) [=Nerinea polymorpha] BR17
‘Cavolinia’ Bruguière 1791 non Abilgaard 1791 BR05
Crenea Risso 1826 BR05
Cypraeogemmula Vredenburg 1920 [Cypraeogemmulinae, Cypraeogemmulini] BR05
`--*C. scabriuscula (Koenen 1890) BR17 [=Trivia scabriuscula BR17, Cypraea scabriuscula H09]
Incrispella Tasch 1963 [Incrispellidae] BR05
`--*I. rectotortis Tasch 1963 BR17
Japeuthria Iredale 1918 [Japeuthriinae] BR05
`--*J. ferrea (Reeve 1847) [=Buccinum ferreum] BR17
Kanamarua Kuroda 1951 [Kanamariidae, Kanamaruidae] BR05
`--*K. adonis (Dall 1919) BR17 [=Colus adonis BR17, C. (Anomalosipho) adonis O27a]
Ladamarekia Horný 1992 [Ladamarekiidae] BR05
`--*L. miranda Horný 1992 BR17
Megalostominae BR05
|--Cataulus BR05
`--Nicida BR05
Melanioptyxis Cossmann 1896 [Melanioptyxinae] BR05
`--*M. altararis (Cossmann 1885) [=Nerinea altararis] BR17
Neoptyxis Wenz 1940 [Neoptyxidae] BR05
`--*N. astrachanica (Rebinder 1902) [=Nerinea astrachanica] BR17
Oligoptyxis Pchelintsev 1953 [Oligoptyxidae] BR05
`--*O. turricula Pchelintsev 1953 BR17
Yuopisthonema Nützel in Bouchet, Rocroi et al. 2017 BR17 (see below for synonymy)
`--*Y. undulatum (Yu 1974) [=*Opisthonema undulatum] BR17
Palaeocyclophorus Wenz 1923 BR05
Pentaptyxis Pchelintsev 1965 [Pentaptyxidae] BR05
`--*P. staszycii (Zeuschner 1849) [=Acteon staszycii] BR17
Plesioplocus Pchelintsev 1953 [Plesioplocidae] BR05
`--*P. grandis Pchelintsev 1953 BR17
Polyodonte Fischer 1807 BR05
Polyodonta Megerle 1811 BR05
Protoconchoididae [Protoconchioididae] BR05
|--Patelliconus Horný 1961 BR17, BR05 [Patelliconidae]
| `--*P. primulus (Perner 1903) [=Palaeacmaea primula] BR17
`--Protoconchoides Shaw 1962 [=Protoconchioides (l. c.)] BR05
|--*P. hermitensis (Resser 1945) [=Scenella hermitensis] BR17
`--P. conula TS11
Pseudobythinella Melville 1956 BR05
Rhytidopilus Cossmann 1895 [=Rhytidophilus (l. c.); Rhytidophilidae, Rhytidopilidae] BR05
`--*R. humbertinus (Buvignier 1852) [=Patella humbertina] KC60
Stylocheilus Gould 1852 [Stylocheilinae, Stylochiniae] BR05
`--*S. lineolatus (Gould 1852) [=Aplysia lineolatus] BR17
Transovula de Gregorio 1880 [Transovulini] BR05
`--*T. schefferi (de Gregorio 1880) [=Ovula schefferi] BR17
Papustyla pulcherrima USDI77
Acilia franciscae TE01
Bathyomphalus contortus (Linnaeus 1758) L06
Caseolus CC06
Leiostyla CC06
Heterostoma Filippi 1837 CC06, LT64 [=Heterostomum Diesing 1850 LT64]
`--H. paupercula (Lowe 1831) [incl. Steenbergia duplex] CC06
Spirorbula CC06
|--S. depauperata (Lowe 1831) [incl. S. latinea (Paiva 1866)] CC06
|--S. obtecta CC06
`--S. squalida CC06
Plagyrona placida (Shuttleworth 1852) CC06
Staurodon saxicola CC06
Actinella CC06
|--A. arcinella CC06
| |--A. a. arcinella CC06
| `--A. a. papillosa CC06
|--A. crassiuscula CC06
|--A. effugiens CC06
|--A. morenensis CC06
`--A. promontoriensis CC06
Pseudocampylaea CC06
|--P. lowei CC06
`--P. portosanctana CC06
Amphorella CC06
|--A. cimensis CC06
|--A. gracilis CC06
|--A. melampoides CC06
|--A. oryza CC06
|--A. terebella CC06
|--A. triticea CC06
`--A. tuberculata CC06
Cylichnidia ovuliformis CC06
Lemniscia michaudi CC06
Eufenella pupoides (Adams 1860) BD86
Clathrofenella reticulata (Adams 1860) BD86
Allocharopa belli SC07
Probaicalia ZZ88
|--P. gerassimovi ZZ88
|--P. rammelmeyeri ZZ88
`--P. vitimensis ZZ88
Brotiopsis wakinoensis ZZ88
Tulotomoides talatziensis ZZ88
Gyrodema etheridgei F71
Magelia bidens Tenison-Woods 1878 F71
Odontostomia cretacea Etheridge 1907 F71
Lindoliva spengleri Petuch 1988 P89
Armiger crista (Linnaeus 1758) P88
Probythinella lacustris (Baker 1928) P88
Bulimnea megasoma (Say 1824) P88
Japonia AA88
|--J. ciliata (Sowerby 1843) AA88
`--J. stephanophora (Moellendorff 1895) AA88
Gymnentome valida H75
Ittibittium Houbrick 1993 BC01
`--I. turriculum (Usticke 1969) (see below for synonymy) BC01
Torrella Henderson & Bartsch 1920 BC01
`--*T. torreiana (Gundlach ex Arango y Molina 1878) [=Ctenopoma torreianum] BC01
Pseudotorinia retifera (Dall 1892) [incl. Heliacus crystallina Usticke 1969] BC01
Bullata Jousseaume 1875 BC01
|--B. matthewsi (van Mol & Tursch 1967) [=Marginella (Prunum) matthewsi] BC01
`--B. ovuliformis WE63
Boonea impressa (Say 1822) [incl. Odostomia (Menestho) beauforti Jacot 1921] BC01
Fargoa bushiana (Bartsch 1909) [incl. Odostomia charlottae Usticke 1969] BC01
Mesophora Laseron 1958 BC01
`--M. novem (Usticke 1969) [=Triphora novem] BC01
Eutriphora bermudensis (Bartsch 1911) [incl. Triphora turrissimilis Usticke 1969] BC01
Aroapyrgus Baker 1931 BC01
`--A. dubiosus (Adams 1852) [=Truncatella dubiosa] BC01
Chippewaella patellitheca W01
Moerkeia Böhm 1895 NE02
Greggelix indigena C90
Torquilla frumentum D56
Sinopupoides hubeiensis NG13
Bulgarica thessalonica (Olivier 1801) O03
Clavilithes rugosus C60
Partulida spiralis C60
Ampullella bulimoides C60
Cepatia cepacea C60
Palaeocyclotus exaratus C60
Pseudorhytidopilus Cox in Knight, Cox et al. 1960 KC60
`--*P. lennieri Cox in Knight, Cox et al. 1960 (see below for synonymy) KC60
Atlantobellerophon Trechmann 1930 KC60
`--*A. zealandicus Trechmann 1930 KC60
Burdikinia Knight 1937 [=Polyamma Etheridge 1917 non Kriechbaumer 1894] KC60
`--*B. burdikiensis (Etheridge 1917) [=*Polyamma burdikiensis] KC60
Cinctaspira Powell 1933 KC60
`--*C. conica Powell 1933 KC60
‘Conchula’ Steininger 1849 non Herrmannsen 1847 (n. d.) KC60
`--*C. cylindracea Steininger 1849 (n. d.) KC60
‘Geinitzia’ Dietz 1911 nec Gemmellaro 1892 nec Handlirsch 1906 KC60
`--*G. carinata Dietz 1911 KC60
Kebina Vologdin 1955 KC60
`--*K. pulchra Vologdin 1955 KC60
Ozarkina Ulrich & Bridge 1931 [=Ozarkispira Ulrich & Bridge 1931 non Walcott 1924] KC60
`--*O. typica (Ulrich & Bridge 1931) [=*Ozarkispira typica] KC60
Ozarkocanus Heller 1956 KC60
`--*O. prearcuatus Heller 1956 KC60
Pondia Oder 1932 KC60
`--*P. powelli Oder 1932 KC60
Pythmenema Lamont & Gilbert 1945 KC60
`--*P. praenantium (Phillips 1848) [=Euomphalus praenantius] KC60
Trochotremaria Ryckholt 1860 KC60
Umbotrochus Perner 1903 [=Umbonitrochus Cossmann 1918] KC60
`--*U. aspersus Perner 1903 KC60
Acevina Rusconi 1952 KC60
`--*A. cuyunchensis (Rusconi 1952) [=Helcionella (*Acevina) cuyunchensis] KC60
Odontosagda V18
|--O. abbotti Vanatta 1918 V18
|--O. blandi V18
`--O. hilli V18
Aplysiopsis smithi (Marcus 1961) B77
Volvaria ficoides H86
Cassidaria H86
|--C. senex [=Struthiolaria senex] H86
`--C. sulcata H86
Sorapisella Bandel 1993 B00
Triadoskenea Bandel 1993 B00
`--T. ampezzana Bandel 1993 B02
Rolandomphalus Schwardt 1992 B00
Macrophragmus tokyoensis K10
Fascioplex neozelanica K95
Pseudoaclisina Yoo 1994 B02
|--*P. turgida [=Aclisina turgida] B02
`--P. microspirulata B02
Palaeoalvania Yoo 1994 B02
`--*P. talenti (Yoo 1994) [=Aclisina talenti] B02
Sinistriconcha Heidelberger & Bandel 1999 B02
|--*S. lierli Heidelberger & Bandel 1999 B02
`--S. minutissima (Yoo 1988) B02
Iberus E99
|--I. dupuydelomei E99
`--I. marmoratus C84
Scabrella Sacco 1890 H22
`--*S. scabra [=Columbella scabra] H22
Pseudaulicina Chavan 1948 BR17
`--*P. musicalis (Lamarck 1803) [=Voluta musicalis, Volutilithes musicalis] BR17
Kimina Yoo 1994 B02
|--*K. globosa Yoo 1994 B02
|--K. australis Yoo 1994 B02
|--K. minor Yoo 1994 B02
`--K. yooi Frýda & Bandel 1997 B02
Harperispira Bandel 2002 BR17
Kiringella Rozov 1968 W01, RJ76
|--K. kultavasaensis RJ76
`--K. pyramidalis W01
Palaeoniso brazieri SS16
Burdekina JB12
Natria JB12
Peraubium JB12
Stusakia pulchra F12
Mesolanistes V-RR93
|--M. bajanohongorensis V-RR93
|--M. debilicostata V-RR93
`--M. shuwalovi V-RR93
Severynella trochlearis Hynda 1983 D94
Lacunina Kittl 1899 TTE93
Bullopsis Conrad 1858 TTE93
Serradonta Okutani, Tsuchida & Fujikara 1992 AJH07
Pleuroprocta silvatica PB27
Trichotoxon PB27
|--T. maculatum PB27
| |--T. m. maculatum PB27
| `--T. m. perforatum PB27
|--T. pardus PB27
`--T. ruwenzoriense PB27
Bocageia PB27
|--B. germaini PB27
|--B. interioris PB27
|--B. liocephala PB27
`--B. runssorina PB27
Africarion PB27
|--A. copiosus PB27
|--A. spatiosus PB27
`--A. tenebrosus PB27
Leucochroopsis E99
|--L. apicalis E99
|--L. dufrenoyi E99
|--L. kleini E99
|--L. leptoloma E99
|--L. pisum E99
`--L. valentinensis E99
|--L. v. valentinensis E99
`--L. v. sermenazensis E99
Wenzia ramondi E99
Megalotachea E99
|--M. christoli E99
|--M. delphinensis E99
| |--M. d. delphinensis E99
| `--M. d. tersannensis E99
|--M. gualinoi E99
`--M. turonensis E99
Tudorella E99
|--T. baudoni E99
`--T. draparnaudi E99
|--T. d. draparnaudi E99
`--T. d. minor E99
Pseudoleacina (Paraglandina) E99
|--P. (P.) christoliana E99
`--P. (P.) oligostropha E99
Pseudochloritis E99
|--P. incrassatus E99
`--P. mollonensis E99
Galactochilus E99
|--G. leobersdorfensis E99
`--G. locardi E99
Negulus E99
|--N. bleicheri E99
|--N. raricostatus E99
|--N. suturalis E99
| |--N. s. suturalis E99
| `--N. s. gracilis E99
|--N. truci E99
`--N. villafranchianus E99
Eostrobilops E99
|--E. aloisii E99
|--E. fischeri E99
`--E. patuliformis E99
Macrozonites E99
|--M. casteti E99
`--M. collongeoni E99
Fortuna seringi E99
Truciella ballesioi E99
Protodrepanostoma E99
|--P. bernardii E99
|--P. nordsiecki E99
`--P. plioauriculatum E99
Monoptychia monoptyx E99
Serruluna anodon E99
Cladopoda H79
|--C. directa H79
`--C. monilifera H79
Carocolla bidentata G40
Clisospiroidea F12
Bellerophontacea [Belleromorpha, Bellerophontina, Fissidorsata, Planspiralia, Prorhipidoglossa] KC60
| i. s.: Patellostium Waagen 1880 (n. d.) KC60
| `--*P. macrostoma (Roemer 1844) [=Bellerophon macrostoma] KC60
| Euphemitella Tasch 1953 (n. d.) KC60
| `--*E. emrichi Tasch 1953 KC60
|--Bellerophontoidea BR17
`--Isospira Koken 1897 KC60 [Isospiridae, Isospiroidea BR05]
`--*I. bucanioides Koken 1897 BR17
Archinacelloidea GS75
Utriculopsis globosa (Lovén 1846) (see below for synonymy) N79
Patelloidea Tasch 1961 T61
`--*P. limnensis Tasch 1961 T61
Permoplanorboidea Tasch 1961 T61
`--*P. primus Tasch 1961 T61
Rugoplanorboidella Tasch 1961 T61
`--*R. sedgwickii Tasch 1961 T61
Wellingtonia Tasch 1961 T61
`--*W. producta Tasch 1961 T61
Tretospira divesuralica G31
Sosiolites vassilievkensis G31
Polyphemopsis G31
|--P. inornata G31
`--P. nitidulum G31
Isognomostoma isognomostoma J63
Platyla polita NB19
|--P. p. polita NB19
`--P. p. regina (Subai 1977) NB19
Paleobulimulus eocenicus Parodiz 1949 P63
Tugurium MP65
|--T. (Tugurium) caribaeum (Petit de la Saussaye 1856) [=Xenophora caribaea] MP65
`--T. longleyi MP65
Sigatica semisulcata (Gray in Beechey 1839) [=Natica semisulcata] MP65
Hemitrochus varians J65
Plagioptycha J65
|--P. gregoriana J65
`--P. macrodon J65
|--P. m. macrodon J65
`--P. m. utowana J65
Bigotella Cossmann 1914 EH19
Acanthonema Grabau in Sherzer & Grabau 1909 BR05 [Acanthonematidae BR17, Acanthonematinae]
Acanthonema newberryi (Meek 1871) TTE93 [=Orthonema newberryi KC60; incl. *A. holopiforme Grabau 1909 KC60, BR17]
Acteonina d’Orbigny 1850 BR05 [=Actaeonina (l. c.) BR05; Acteoninidae, Acteonininae, Acteoninoidea BR17]
Apogastropoda [Anaclodonta, Architectonicoida, Chalazaeata, Ctenobranchia, Davisianoidei, Dioecia, Discopoda, Ectobranchia, Entobranchia, Entomotaeniata, Gymnoglossa, Heterogastropoda, Heterostropha, Holonephridia, Homoestropha, Hydrocenoidei, Iniophthalma, Loxonematacea, Monopleurobranchia, Monotocardia, Murchisoniata, Musioglossata, Oligopteria, Orbacea, Paludinimorpha, Pectinibranchia, Pectinibranchiata, Peristomacea, Peristomania, Peristomidae, Pneumonobranchiata, Potamophila, Proboscidifera, Procaenogastropoda, Ptenobranchiata, Pyramidellida, Pyramidellimorpha, Pyramidelliones, Pyramidelloida, Rissoellina, Solenisciformes, Spartaebranchia, Subulitiformes, Taenioglossa, Trachelobranchia, Volumina] S11
Caenogastropoda [Architaenioglossa, Architaenioglossae, Basiophthalma, Cyclophoracea, Cyclophoroidei, Fusiformia, Globularioidei, Mesogastropoda, Mesogastropodida, Opisthophthalma, Palaeocyclophoridae, Procyclophoridae, Procyclophoroidea, Rostrifera, Teleogeophila] S11
Dendropupa Owen 1859 BR17 [incl. Strophella Dawson 1895 KC60, Strophites Dawson 1880 non Deshayes 1832 KC60; Dendropupidae]
Gastropoda [Abranchiata, Alata, Anentomostomata, Anisobranchia, Anisopleura, Aponotoneura, Apterygia, Archaeogastropoda, Archaeoprosobranchia, Arthrocochlides, Asiphonobranchia, Asiphonata, Aspidobranchia, Auxogastropoda, Chiastoneura, Chismobranchiata, Clypeaceae, Cocculinacea, Cocculiniformia, Cochliostraca, Conivalvia, Cricostomata, Ctenobranchiata, Cyclobranchia, Dicoeca, Dicranobranchia, Diotocardia, Eogastropoda, Fissobranchia, Fissuracea, Fornices, Galeroconcha, Gasteropoda, Gasteropodophora, Gasterozoa, Gastraeopoda, Gastropodea, Goniostomata, Gonyostomata, Gymnopoda, Helcionellina, Hemicyclostoma, Holostomata, Hyperstrophina, Lepadophora, Lepetellina, Limaces, Neurobranchia, Operculata, Orthogastropoda, Otidea, Paracephalophora, Paragastropoda, Patellina, Peristomatacea, Peristomatidae, Platyceratacea, Podophthalma, Prosobranchia, Prosobranchiata, Pselaphocephala, Raphidoglossa, Rhipidoglossa, Riphidoglossa, Scutiformia, Sinuata, Sinuopeoidei, Solenostomata, Streptoneura, Syncephala, Tectipeda, Triforidoidei, Trochacea, Trochina, Trochomorpha, Trochomorphi, Turbinimorpha, Xenophoroidei]
Ittibittium turriculum (Usticke 1969) [=Bittium varium f. turricula, Alabina turricula, B. turriculum, Cerithiodes turricula] BC01
Kittlidiscus Haas 1953 BR05 [=Schizodiscus Kittl 1891 non Hall & Clarke 1888 KC60; Kittlidiscidae BR17]
*Kittlidiscus bronni (Klipstein 1845) KC60 [=Pleurotomaria bronni KC60, Schizodiscus bronni KC60; incl. P. plana Klipstein 1845 non Münster 1844 KC60, *Kittlidiscus planus BR17, *Schizodiscus planus BR17]
*Pseudorhytidopilus lennieri Cox in Knight, Cox et al. 1960 [incl. Helcion castellana Lennier 1868 non Patella castellana Thurmann & Étallon 1861] KC60
Sorbeoconcha [Anandria, Aulobranchiata, Brevicommisurata, Eriophthalma, Neotaenioglossa, Omalaxoidea, Palaeocaenogastropoda, Siphonobranchei, Siphonobranchia, Siphonobranchiata, Tubispirantia] S11
Spirostylus linctus (Böhm 1895) [=Omphaloptycha lincta; incl. O. porrecta Böhm 1895] TE01
Utriculopsis globosa (Lovén 1846) [=Amphisphyra globosa, Utriculus globosus; incl. Utriculopsis vitrea Sars 1870] N79
Yuopisthonema Nützel in Bouchet, Rocroi et al. 2017 BR17 [=Opisthonema Yu 1974 non Gill 1862 BR05; Opisthonematidae, Yuopisthonematidae]
*Type species of generic name indicated
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