Lagenida

Longitudinal section of Syzrania pulchra, from Groves et al. (2003).

Belongs within: Foraminifera.
Contains: Nodosariidae, Polymorphinidae, Glandulinidae, Geinitzinidae.

The Lagenida are a group of foraminifera with a low-magnesium, calcitic secreted test that is monolamellar and radial (Groves et al. 2004) They first appeared in the middle late Carboniferous, though they did not become prominent until the Middle Permian (Grobes et al. 2003).

<==Lagenida [Lagenina, Nodosariata, Syzraniidae]
    |--Syzrania Reitlinger 1950 GAR03
    |    |--*S. bella Reitlinger 1950 [=Earlandia bella] LT64
    |    |--S. amazonica Atliner & Savini 1997 GAR03
    |    |--S. grovesi Altiner & Savini 1997 GAR03
    |    `--S. pulchra Kireeva 1958 GAR03
    `--+--Tezaquina Vachard 1980 [incl. Syzranella] GAR03
       |    `--*T. clivuli Vachard 1980 GAR03
       |--Amphoratheca Mamet & Pinard 1992 GAR03
       |    `--A. iniqua Mamet & Pinard 1992 GAR03
       `--+--Geinitzinidae GAR03
          `--Protonodosariidae GAR03
               |--Frondina Sellier de Civrieux & Dessauvagie 1965 GRA04
               |--Tauridia Sellier de Civrieux & Dessauvagie 1965 GRA04
               `--Protonodosaria Gerke 1959 GAR03
                    |--*P. proceraformis (Gerke 1952) GRA04 [=Nodosaria proceraformis LT64]
                    |--P. praecursor (Rauser-Chernousva 1949) GRA04
                    `--P. rauserae Gerke 1959 GAR03

Lagenida incertae sedis:
  Polarisella Mamet & Pinard 1992 GAR03
    `--*P. blindensis Mamet & Pinard 1992 GAR03
  Nodosariacea [Enclinostegia, Lagenicae, Lagenidea, Nodosalidia, Titanostichostegia] S-VC91
    |--Nodosariidae LT64
    |--Polymorphinidae LT64
    `--Glandulinidae S-VC91
  Nodosinellidae [Nodosinellida, Nodosinellinae] LT64
    |--Frondilina Bykova 1952 LT64
    |    `--*F. devexis Bykova 1952 LT64
    |--Hipporina Bykova in Bykova & Polenova 1955 [=Hypporina (l. c.)] LT64
    |    `--*H. hastila Bykova in Bykova & Polenova 1955 LT64
    |--Lunucammina Spandel 1898 (n. d.) (see below for synonymy) LT64
    |    |--*L. permiana (Spandel 1898) [=Geinitzella (*Lunucammina) permiana] LT64
    |    |--‘*Eogeinitzina’ devonica Lipina 1950 LT64
    |    |--L. excavata (Cushman & Waters 1928) [=*Spandelina excavata] LT64
    |    |--L. orientalis (Miklukho-Maklay 1954) [=*Neogeinitzina orientalis] LT64
    |    `--L. perforata (Lange 1925) [=*Padangia perforata] LT64
    `--Nodosinella Brady 1876 (n. d.) GRA04 (see below for synonymy)
         |--*N. digitata Brady 1876 GRA04 [=*Arnodosinum digitatum LT64]
         |--‘*Monogenerina’ atava Spandel 1901 LT64
         |--‘*Eonodosaria’ evlanensis Lipina 1950 LT64
         |--N. glennensis C40
         |--N. measpis (Bykova 1952) [=*Tikhinella measpis] LT64
         `--N. nodosariformis (Cushman & Waters 1928) [=Spandelina (*Spandelinoides) nodosariformis] LT64
  Olympina Reichel 1946 GRA04
    `--*O. insolita Reichel 1945 LT64
  Colaniellidae [Colaniellinae] GRA04
    |--Multiseptida Bykova 1952 LT64
    |    `--*M. corallina Bykova 1952 LT64
    `--Colaniella Licharev 1939 W76 (see below for synonymy)
         |--*C. parva (Colani 1924) [=*Pyramis parva] LT64
         |--C. cylindrica M.-Maclay 1954 W76
         |--C. nana M.-Maclay 1954 W76
         `--C. ussuriensis (Sosnina in Kiparisova et al. 1956) [=*Wanganella ussuriensis] LT64

Colaniella Licharev 1939 W76 [=Pyramis Colani 1924 nec Röding 1798 nec Schumacher 1817 LT64; incl. Wanganella Sosnina in Kiparisova et al. 1956 LT64]

Lunucammina Spandel 1898 (n. d.) [incl. Eogeinitzina Lipina 1950, Neogeinitzina Miklukho-Maklay 1954, Padangia Lange 1925 nec Babor 1900 nec Werner 1924, Spandelina Cushman & Waters 1928] LT64

Nodosinella Brady 1876 (n. d.) GRA04 [=Arnodosinum Rhumbler 1913 LT64; incl. Eonodosaria Lipina 1950 LT64, Monogenerina Spandel 1901 LT64, Spandelinoides Cushman & Waters 1928 LT64, Tikhinella Bykova 1952 LT64]

*Type species of generic name indicated

REFERENCES

[C40] Cushman, J. A. 1940. Foraminifera: Their classification and economic use 3^rd ed. Harvard University Press: Cambridge (Massachusetts).

[GAR03] Groves, J. R., D. Altiner & R. Rettori. 2003. Origin and early evolutionary radiation of the order Lagenida (Foraminifera). Journal of Paleontology 77 (5): 831–843.

[GRA04] Groves, J. R., R. Rettori & D. Altiner. 2004. Wall structures in selected Paleozoic lagenide Foraminifera. Journal of Paleontology 78: 245–256.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[S-VC91] Segura-Vernis, L. R., & A. L. Carreño. 1991. Foraminíferos y ostrácodos de la Laguna de La Paz, Baja California Sur, México. Inv. Mar. CICIMAR 6 (1): 195–224.

[W76] Wang K. 1976. The foraminifera from the Changhsing Formation in western Guizhou. Acta Palaeontologica Sinica 15 (2): 187–195.