Chordeumatida

Specimen belonging to the Australeuma mauriesi group, from Tasmanian Multipedes.

Belongs within: Eugnatha.
Contains: Neoatractosomatoidea, Cleidogonoidea, Metopidiotrichinae, Conotylidae.

The Chordeumatida are a group of relatively short, bristly millipedes.

Characters (from Hoffman 1990): Body with 26–30 segments in adults; ocelli usually present; gnathochilarium with short basal mentum and elongate lingual lamellae; ozopores absent; metaterga with well-defined median longitudinal suture; sterna not firmly attached to adjoining sternites but more or less flexibly articulated by connective tissue; males with coxal sacs in ninth and tenth pairs of legs only; gonopods composed of both pairs of legs of seventh segment.

<==Chordeumatida [Chordeumida]
    |--Chordeumatidea S03b
    |    |  i. s.: Chordeumella [Chordeumatidae] M03
    |    |           `--C. broelemanni (Verhoeff 1897) M03
    |    `--Cleidogonoidea H90
    |--Striariidea S03b
    |    |--Caseyidae [Caseyoidea] S03b
    |    |    |--Caseya H90
    |    |    |--Opiona H90
    |    |    |--Placerna H90
    |    |    |--Vasingtona H90
    |    |    |--Zantona H90
    |    |    `--Underwoodia H90
    |    `--Striarioidea S03b
    |         |--Apterourus Loomis 1966 [Apterouridae] S03b
    |         |    |--*A. horizontalis Loomis 1966 S03b
    |         |    `--A. serratus Shear 2003 S03b
    |         `--+--Rhiscosomides H90 [Rhiscosomididae S03b]
    |            `--Striariidae S03b
    |                 |--Vaferia H90
    |                 `--Speostriaria H90
    |--Brannerioidea [Craspedosmatidea] SL03
    |    |--Niponiosoma [Niponiosomatidae] SH98
    |    |--Trachygonidae SL03
    |    |--Heterolatzeliidae SL03
    |    |--Chaemosomatidae SL03
    |    |--Microlympia Shear & Leonard 2003 [Microlympiidae] SL03
    |    |    `--*M. echina Shear & Leonard 2003 SL03
    |    |--Branneria Bollman 1893 [Branneriidae] S03a
    |    |    |--*B. carinata (Bollman 1888) [=Craspedosoma carinata] S03a
    |    |    `--B. bonoculus Shear 2003 S03a
    |    |--Tingupidae SL03
    |    |    |--Tingupa Chamberlin 1910 SL03
    |    |    |    `--T. pallida SH98
    |    |    |--Buotus Chamberlin 1940 SL03
    |    |    |    `--B. carolinus SH98
    |    |    `--Blancosoma Shear & Hubbard 1998 SL03
    |    |         `--*B. scaturgo Shear & Hubbard 1998 SH98
    |    `--Kashmireumatidae S02b
    |         |--Kashmireuma S02b
    |         |    |--K. nepalensis S02b
    |         |    |--K. nielseni S02b
    |         |    `--K. schawallari S02b
    |         `--Vieteuma Golovatch 1984 S02b
    |              |--*V. topali S02b
    |              |--V. hubeiensis Mauriés & Nguyen-Duy Jaquemin 1997 S02b
    |              `--V. longi Shear 2002 S02b
    `--Heterochordeumatoidea [Heterochordeumatidea] S03b
         |--Heterochordeumatidae S02a
         |--Metopidiotrichidae [Metopidiothricidae] S02a
         |    |--Schedotrigona Silvestri 1903 [Schedotrigonidae, Schedotrigoninae] S02a
         |    |--Metopidiotrichinae S02a
         |    `--Neocambrisomatinae [Neocambrisomatidae] S02a
         |         |--Neocambrisoma Mauriès 1987 S02a
         |         |--Nesiothrix Shear & Mesibov 1997 S02a
         |         |--Nipponothrix Shear & Tanabe 1994 S02a
         |         `--Australeuma Golovatch 1987 S02a
         |              `--A. mauriesi Shear & Mesibov 1997 S02a
         |--Megalotylidae S02b
         |    |--Megalotyla S02b
         |    `--Nepalella Shear 1987 S02b
         |         |--N. birmanica S02b
         |         |--N. caeca Shear 1999 S02b
         |         |--N. griswoldi Shear 2002 S02b
         |         |--N. kavanaughi Shear 2002 S02b
         |         |--N. magna Shear 2002 S02b
         |         `--N. pianma Shear 2002 S02b
         |--Adrityla [Adritylidae] H90
         `--Conotylidae H90

Chordeumatida incertae sedis:
  Diplomaragna [Diplomaragnidae] S03a
  Neoatractosomatoidea M03
  Mastigophorophylloidea M03
    |--Mastigophorophyllidae M03
    |--Hoffmaneumatidae M03
    `--Altajellidae M03
  Anthroleucosomatidae [Antroleucosomatoidea] M03
    |--Krueperia nivale Verhoeff 1900 M03
    `--Anamastigona M03
         |--A. bilselii Verhoeff 1940 M03
         |--A. hauseri Strasser 1974 M03
         |--A. matsakisi Mauriès & Karamaouna 1984 M03
         |--A. mediterranea Curčić, Makarov & Lymberakis 2001 M03
         |--A. penicillata Attems 1902 M03
         `--A. pentelonica (Verhoeff 1925) M03
  Nanogona Cook 1895 H00
  Urochordeuma [Urochordeumatidae] H90

*Type species of generic name indicated

REFERENCES

[H00] Hannibal, J. T. 2000. Hexecontasoma, a new helminthomorph millipede (Hexecontasomatidae n. fam.) from the Mazon Creek, Illinois, fauna (Carboniferous, North America). Fragmenta Faunistica 43 (Suppl): 19-35.

[H90] Hoffman, R. L. 1990. Diplopoda. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 835-860. John Wiley & Sones: New York.

[M03] Mauriès, J.-P. 2003. Schizmohetera olympica sp. n. from Greece, with a reclassification of the superfamily Neoatractosomatoidea (Diplopoda: Chordeumatida). Arthropoda Selecta 12: 9-16.

[S02a] Shear, W. A. 2002a. The millipede genus Metopidiothrix Attems (Diplopoda: Chordeumatida: Metopidiotrichidae). Invertebrate Systematics 16: 849-892.

[S02b] Shear, W. A. 2002b. Five new chordeumatidan millipeds from China: New species of Vieteuma (Kashmireumatidae) and Nepalella (Megalotylidae). Proceedings of the California Academy of Sciences 53 (6): 63-72..

[S03a] Shear, W. A. 2003a. Branneria bonoculus, n. sp., a second species in the North American milliped family Branneriidae (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa 233: 1-7.

[S03b] Shear, W. A. 2003b. A second species of the rare milliped family Apterouridae (Diplopoda: Chordeumatida: Striarioidea). Proceedings of the Biological Society of Washington 116 (2): 509-514.

[SH98] Shear, W. A. & D. A. Hubbard, Jr. 1998. Cave millipeds of the United States. IV. A new genus and species from a high altitude cave in Colorado (Diplopoda, Chordeumatida, Tingupidae). Myriapodologica 5 (8): 85-94.

[SL03] Shear, W. A., & W. P. Leonard. 2003. Microlympiidae, a new milliped family from North America, and Microlympia echina, new genus and species (Diplopoda: Chordeumatida: Brannerioidea). Zootaxa 243: 1-11.