Loeblichiidae

Axial (left) and equatorial sections of Lysella gadukensis, from Cózar & Vachard (2001).

Belongs within: Fusulinida.
Contains: Dainella, Bessiella, Paralysella, Ozawainellidae, Paradainella, Loeblichiinae, Staffellidae, Schubertellidae, Verbeekinidae, Fusulininae, Schwagerininae.

The Loeblichiidae are a lineage of fusulinid Foraminifera from the lower Carboniferous with numerous chambers and true chomata that were regarded by Cózar & Vachard (2001) as including the ancestors of later fusulinids from the later Carboniferous and Permian. The latter, comprising the Fusulinacea, developed a distinctively complex internal structure, with the interiors of the numerous, short chambers modified by secondary deposits such as chomata, parachomata, tectoria or axial fillings (Loeblich & Tappan 1964).

<==Loeblichiidae [Loeblichinidae, Spinoendothyrinae]
    |--Inflatoendothyra Brazhnikova & Vdovenko in Vdovenko 1972 CV01
    |--Elergella Conil in Shilo et al. 1984 CV01
    `--+--Spinoendothyra Lipina in Anonymous 1963 CV01
       |--+--Spinobrunsiina Conil & Longerstaey in Conil et al. 1980 CV01
       |  `--Pseudoplanoendothyra Brazhnikova & Vdovenko in Brazhnikova 1982 CV01
       |--+--Urbanella Malakhova in Anonymous 1963 CV01
       |  `--Banffella Mamet in Mamet & Skipp 1970 CV01
       `--+--Pseudochernyshinella Brazhnikova 1974 CV01
          `--Dainellinae CV01
               |--Dainella CV01
               |--+--Bessiella CV01
               |  `--Vissarionovella Cózar & Vachard 2001 CV01
               |       |--*V. tujmasensis (Vissarionova 1948) (see below for synonymy) CV01
               |       |--V. donzellii Cózar & Vachard 2001 CV01
               |       `--V. llangollensis (Conil & Ramsbottom in Conil et al. 1981) [=Bessiella llangollensis] CV01
               |--+--Lysella Bozorgnia 1973 CV01
               |  |    |--*L. gadukensis Bozorgnia 1973 [incl. L. conferta Bozorgnia 1973] CV01
               |  |    |--L. amenta (Ganelina 1966) [=Dainella amenta] CV01
               |  |    |--L. compacta (Postoyalko 1970) [=Dainella compacta] CV01
               |  |    |--L. incomposita (Postoyalko 1970) [=Dainella incomposita] CV01
               |  |    |--L. magna (Vdovenko 1954) [=Quasiendothyra magna] CV01
               |  |    |--L. manifesta (Ganelina 1966) [=Dainella manifesta] CV01
               |  |    |--L. mediocriformis CV01
               |  |    |--L. micula (Postoyalko 1970) [=Dainella micula] CV01
               |  |    `--L. uralica (Postoyalko 1970) [=Dainella uralica] CV01
               |  `--+--Paralysella CV01
               |     `--+--Ozawainellidae CV01
               |        `--Eostaffella Rauzer-Chernousova 1948 CV01, LT64 [Eostaffellidae]
               |             |--*E. parastruvei (Rauzer-Chernousova 1948) [=Staffella (*Eostaffella) parastruvei] LT64
               |             |--E. compacta Rozovskaya 1963 [=E. versabilis compacta] CV01
               |             |--E. endothyroidea Chang 1962 CV01
               |             |--E. nalivkini Malakhova 1957 CV01
               |             |--E. prisca CV01
               |             |    |--E. p. prisca CV01
               |             |    `--E. p. ocisa Vdovenko 1971 CV01
               |             `--E. radiata CM01
               `--+--Paradainella CV01
                  |--+--Eogloboendothyra CV01
                  |  `--+--Skippella CV01
                  |     `--Eoendothyranopsis CV01
                  |          |--E. macrus R79
                  |          |--E. rarus R79
                  |          |--E. spiroides R79
                  |          `--E. utahensis R79
                  `--+--Loeblichiinae CV01
                     |--Bozorgniella Cózar & Vachard 2001 CV01
                     |    |--*B. tumultuosa (Bozorgnia 1973) [=Dainella tumultuosa] CV01
                     |    |--B. florigena (Pronina 1963) [=Pseudoendothyra florigena, Eoparastaffella florigena] CV01
                     |    |--B. fundata (Simonova 1975) [=Eoparastaffella fundata] CV01
                     |    |--B. librans (Conil & Lys 1964) [=Plectogyra librans] CV01
                     |    |--B. rotunda (Vdovenko 1971) [=Eoparastaffella (Eoparastaffellina) rotunda] CV01
                     |    |--B. simplex (Vdovenko 1954) [=Eoparastaffella simplex] CV01
                     |    `--B. tumida (Pronina 1963) [=Eoparastaffella tumida, Pseudoendothyra tumida] CV01
                     `--+--Eoparastaffelina umbilicata (Bozorgnia 1973) [=Lysella umbilicata] CV01
                        `--Fusulinacea (see below for synonymy) K04
                             |  i. s.: Chenella Miklukho-Maklay 1959 LT64
                             |           `--*C. kueichihensis (Chen 1934) [=Orobias kueichihensis] LT64
                             |         Lantschichites Tumanskaya 1953 LT64
                             |           `--*L. maslennikovi (Tumanskaya 1953) (see below for synonymy) LT64
                             |         Darvasites Miklukho-Maklay 1959 LT64
                             |           `--*D. daroni (Miklukho-Maklay 1949) (see below for synonymy) LT64
                             |         Eostaffelloides Miklukho-Maklay 1959 LT64
                             |           `--*E. orientalis Miklukho-Maklay 1959 LT64
                             |         Fujimotoella Morikawa 1952 LT64
                             |           `--*F. umblicata Morikawa 1952 LT64
                             |         Moscoviella Miklukho-Maklay 1952 [=Moscovella Thompson 1954] LT64
                             |           `--*M. mosquensis (Rauzer-Chernousova in Rauzer-Chernousova et al. 1951) (see below for synonymy) LT64
                             |         Parareichelina Miklukho-Maklay in Rauzer-Chernousova & Fursenko 1959 LT64
                             |           `--*P. reticulata Miklukho-Maklay in Rauzer-Chernousova & Fursenko 1959 LT64
                             |         Praeparafusulina Tumanskaya 1962 LT64
                             |           `--*P. pseudojaponica (Dutkevitch in Gorsky 1939) (see below for synonymy) LT64
                             |         Pseudoendothyra Mikhaylov 1939 (see below for synonymy) LT64
                             |           |--*P. struvii (von Möller 1879) (see below for synonymy) LT64
                             |           `--P. kremenskensis R79
                             |         Putrella Rauzer-Chernousova 1951 LT64
                             |           `--*P. brazhnikovae (Putrya 1948) [=Pseudotriticites brazhnikovae] LT64
                             |         Quasifusulinoides Rauzer-Chernousova & Rozovskaya in Rauzer-Chernousova & Fursenko 1959 LT64
                             |           `--*Q. fusiformis (Rozovskaya 1952) [=Pseudotriticites fusiformis] LT64
                             |         Russiella Miklukho-Maklay 1957 LT64
                             |           `--*R. pulchra Miklukho-Maklay 1957 LT64
                             |         Sichotenella Tumanskaya 1953 LT64
                             |           `--*S. sutschanica Tumanskaya 1953 LT64
                             |         Sphaeroschwagerina Miklukho-Maklay 1959 LT64
                             |           |--S. karnica (Shcherbovich in Rauzer-Chernousova & Shcherbovich 1949) (see below for synonymy) LT64
                             |           |--S. fusiformis RR79
                             |           |--S. moelleri RR79
                             |           |--S. sphaerica RR79
                             |           `--S. vulgaris RR79
                             |--Staffellidae LT64
                             |--Schubertellidae K04
                             |--Verbeekinidae LT64
                             `--Fusulinidae [Fusilinidae, Schwagerinidae] CV01
                                  |--Fusulininae LT64
                                  `--Schwagerininae LT64

*Darvasites daroni (Miklukho-Maklay 1949) [=Triticites ordinatus var. daroni] LT64

Fusulinacea [Fusulinaceae, Fusulinidea, Fusulinoidea, Neoschwagerinaceae, Verbeekinacea, Verbeekinidea] K04

*Lantschichites maslennikovi (Tumanskaya 1953) [=Codonofusiella (*Lantschichites) maslennikovi] LT64

*Moscoviella mosquensis (Rauzer-Chernousova in Rauzer-Chernousova et al. 1951) [=Ozawainella mosquensis, *Moscovella mosquensis] LT64

*Praeparafusulina pseudojaponica (Dutkevitch in Gorsky 1939) [=Parafusulina pseudojaponica] LT64

Pseudoendothyra Mikhaylov 1939 [=Parastaffella Rauzer-Chernousova 1948; incl. Eoparastaffella Vdovenko 1954] LT64

*Pseudoendothyra struvii (von Möller 1879) [=Fusulinella struvii, *Parastaffella struvii] LT64

Sphaeroschwagerina karnica (Shcherbovich in Rauzer-Chernousova & Shcherbovich 1949) [=Schwagerina sphaerica var. karnica; incl. Sc. princeps Schellwien 1898 non Borelis princeps Ehrenberg 1842, *Sphaeroschwagerina princeps] LT64

*Vissarionovella tujmasensis (Vissarionova 1948) [=Eostaffella tujmasensis, Dainella tujmasensis, Lysella tujmasensis] CV01

*Type species of generic name indicated

REFERENCES

[CM01] Cózar, P., & B. L. Mamet. 2001. Planohowchinia (Lasiodiscidae, Foraminiferida), a new Late Viséan genus, southwestern Spain. Journal of Foraminiferal Research 31 (3): 228–232.

[CV01] Cózar, P., & D. Vachard. 2001. Dainellinae subfam. nov. (Foraminiferida du Carbomifère inférieur), révision et nouveaux taxons. Geobios 34 (5): 505–526.

[K04] Kobayashi, F. 2004. Late Permian foraminifers from the Limestone Block in the Southern Chichibu Terrane of west Shikoku, SW Japan. Journal of Paleontology 78 (1): 62–70.

[LT64] Loeblich, A. R., Jr & H. Tappan. 1964. Sarcodina: chiefly “thecamoebians” and Foraminiferida. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt C. Protista 2 vol. 1. The Geological Society of America and The University of Kansas Press.

[R79] Ross, C. A. 1979. Carboniferous. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A254–A290. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).

[RR79] Ross, C. A., & J. R. P. Ross. 1979. Permian. In: Robison, R. A., & C. Teichert (eds) Treatise on Invertebrate Paleontology pt A. Introduction. Fossilisation (Taphonomy), Biogeography and Biostratigraphy pp. A291–A350. The Geological Society of America, Inc.: Boulder (Colorado), and The University of Kansas: Lawrence (Kansas).