Leptoceridae

Notalina sp., copyright Ken Harris.

Belongs within: Leptoceroidea.
Contains: Oecetis, Triplectides.

The Leptoceridae, long-horned caddisflies, are a cosmopolitan group of caddisflies with long, slender wings and notably long, filiform antennae.

<==Leptoceridae
    |  i. s.: Erotesis IS02
    |         Oligoplectrum IS02
    |--Leptocerinae BZ-M04
    |    |  i. s.: Leptocerus Leach 1815 N83
    |    |           |--*L. interruptus (Fabricius 1775) [=Phryganea interrupta] N83
    |    |           |--L. assimulans (Ulmer 1916) [=Setodes assimulans] N83
    |    |           `--L. scuta Mosely 1953 N83
    |    |         Leptorussa Mosely 1953 N83
    |    |           `--*L. darlingtoni (Banks 1939) [=Leptocerus darlingtoni; incl. Leptorussa russata Mosely 1953] N83
    |    |         Oecetina Banks 1899 N83
    |    |         Nectopsyche candida KP19
    |    |--Oecetis BZ-M04
    |    |--Mystacides Berthold 1827 [Mysacidini] BZ-M04
    |    |    `--M. azurea (Linnaeus 1761) BZ-M04
    |    |--Setodes Rambur 1842 [Setodini] BZ-M04
    |    |    |--*S. viridis (Fourcroy 1785) [=Phryganea viridis] N83
    |    |    |--S. argentipunctellus McLachlan 1877 BZ-M04
    |    |    `--S. portionalis S02
    |    |--Athripsodini BZ-M04
    |    |    |--Ceraclea Stephens 1829 BZ-M04
    |    |    |    `--C. sobradieli (Navás 1917) BZ-M04
    |    |    `--Athripsodes Billberg 1820 BZ-M04
    |    |         |--A. albifrons (Linnaeus 1758) BZ-M04
    |    |         `--A. taounate BZ-M04
    |    `--Triaenodini BZ-M04
    |         |--Adicella McLachlan 1877 BZ-M04
    |         |    `--A. reducta (McLachlan 1865) BZ-M04
    |         `--Triaenodes McLachlan 1865 N83
    |              |--*T. bicolor (Curtis 1834) [=Leptocerus bicolor] N83
    |              |--T. bernaysae Korboot 1964 N83
    |              |--T. detruncatus Martynov 1924 T76
    |              |--T. intricata Neboiss 1977 N83
    |              |--T. jubatus Neboiss 1982 N83
    |              |--T. reuteri T76
    |              |--T. simulans Tjeder 1929 [incl. T. forsslundi Tjeder 1941] T76
    |              `--T. volda Mosely 1953 N83
    `--Triplectidinae N83
         |--Triplectides N83
         |--Hudsonema Mosely 1963 N83
         |--Loticana Mosely 1936 N83
         |--Notanatolica McLachlan 1866 N83
         |--Triplexina Mosely 1953 N83
         |--Triplexa Mosely 1953 N83
         |    `--*T. villa Mosely 1953 N83
         |--Condocerus Neboiss 1977 N83
         |    |--*C. paludosus Neboiss 1977 N83
         |    `--C. aptus Neboiss 1982 N83
         |--Lectrides Mosely 1953 N83
         |    |--*L. varians Mosely 1953 N83
         |    `--L. parilis Neboiss 1982 N83
         |--Triplectidina Mosely 1936 N83
         |    |--*T. oreolimnetes (Tillyard 1924) [=Triplectides oreolimnetes] N83
         |    `--T. nigricornis Mosely 1936 N83
         |--Symphitoneuria Ulmer 1906 N83
         |    |--*S. exigua (McLachlan 1862) [=Leptocerus exiguus] N83
         |    |--S. opposita (Walker 1852) [=Leptocerus oppositus, Loticana opposita, Notanatolica opposita] N83
         |    `--S. wheeleri Banks 1939 N83
         |--Westriplectes Neboiss 1977 N83
         |    |--*W. pedderensis Neboiss 1977 N83
         |    |--W. albanus (Mosely 1953) [=Triplectides albana] N83
         |    `--W. angelae Neboiss 1978 N83
         |--Notoperata Neboiss 1977 N83
         |    |--*N. sparsa (Kimmins 1953) [=Hudsonema sparsa] N83
         |    |--N. diversa Neboiss 1982 N83
         |    |--N. maculata (Mosely 1953) [=Hudsonema maculata; incl. Triplexina lobata Jacquemart 1965] N83
         |    |--N. syncope Neboiss 1982 N83
         |    `--N. tenax Neboiss 1982 N83
         `--Notalina Mosely 1936 N83
              |--*N. parkeri Mosely 1936 N83
              |--N. bifaria Neboiss 1977 N83
              |--N. delicatula (Ulmer 1908) [=Triplectides delicatula] N83
              |--N. dwellinga Neboiss 1982 N83
              |--N. flava (Ulmer 1908) [=Triplectides flava] N83
              |--N. fulva Kimmins 1953 N83
              |--N. moselyi Kimmins 1953 N83
              |--N. nigra (Mosely 1953) [=Triplexina nigra] N83
              |--N. pseudodelicata Neboiss 1982 N83
              `--N. tillyardi Kimmins 1953 N83

*Type species of generic name indicated

REFERENCES

[BZ-M04] Bonada, N., C. Zamora-Muñoz, M. Rieradevall & N. Prat. 2004. Trichoptera (Insecta) collected in Mediterranean river basins of the Iberian Peninsula: taxonomic remarks and notes on ecology. Graellsia 60 (1): 41–69.

[IS02] Ivanov, V. D., & I. D. Sukatsheva. 2002. Order Trichoptera Kirby, 1813. The caddisflies (=Phryganeida Latreille, 1810). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 199–220. Kluwer Academic Publishers: Dordrecht.

[KP19] Kawahara, A. Y., D. Plotkin, M. Espeland, K. Meusemann, E. F. A. Toussaint, A. Donath, F. Gimnich, P. B. Frandsen, A. Zwick, M. dos Reis, J. R. Barber, R. S. Peters, S. Liu, X. Zhou, C. Mayer, L. Podsiadlowski, C. Storer, J. E. Yack, B. Misof & J. W. Breinholt. 2019. Phylogenomics reveals the evolutionary timing and pattern of butterflies and moths. Proceedings of the National Academy of Sciences of the USA 116 (45): 22657–22663.

[N83] Neboiss, A. 1983. Checklist and bibliography of the Australian caddis-flies (Trichoptera). Australian Society for Limnology Special Publication 5: 1–132.

[S02] Sinitshenkova, N. D. 2002. Ecological history of the aquatic insects. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 388–426. Kluwer Academic Publishers: Dordrecht.

[T76] Tobias, W. 1976. Köcherfliegen und Steinfliegen einiger Gewässer in Sör Varanger (Nord-Norwegen) (Trichoptera, Plecoptera). IV. Triaenodes detruncatus Martynov neu für Norwegen. Senckenbergiana Biologica 57 (1–3): 49–53.