Belongs within: Cucujoidea.
The Cryptophilinae are a group of saprophagous or mycophagous beetles mostly found amongst leaf litter (Leschen 2003).
Characters (from Leschen 2003): Mandible usually with subapical serrations; lacinia with one or no lacinial spines; mentum lacking lateral pockets, usually with medial carina; apical labial palpomere as wide as or wider than long, usually not securiform; ligular membrane as wide or wider than prementum; transverse gular line usually present as shallow groove (rarely absent or deep); antennal insertion hidden in dorsal view; supraocular line usually present to level above eye; vertex of head commonly with stridulatory file; pronotum usually not strongly constricted at base; anterior angles of pronotum poorly or well developed; anterior margin of pronotum usually serrate; pronotal pits usually present; procoxal cavity internally closed, usually open behind or closed by hypomeron, edge of hypomeron usually smooth; mesometaventral articulation, if present, dicondylic; width of mesoventral process usually less than coxa; submesocoxal lines usually absent; abdominal ventrites 1 and 2 usually free; abdominal calli present; metafurcal lamina usually absent with median stalk of metendosternite poorly or well developed; spermatheca rounded or elongate; apical pit of spermatheca usually absent; accessory gland of spermatheca usually present; tarsomere 1 usually longer than tarsomere 2 and 3; tarsomere 4 usually exposed in ventral view; tarsal shelf of tarsomere 5 usually absent; empodium with one or two setae; elytral punctation striate or confused with scutellary striole absent; elytra usually narrowly explanate; hind with with radial cell and wedge cell absent.
<==Cryptophilinae [Cryptophilidae]
|--Empocryptini L03
| |--Lobosternum clavicorna L03
| `--+--Lepidotoramus grouvellei L03
| `--Empocryptus Sharp 1900 L03
| `--E. ovalis L03
`--+--Toramini [Toraminae] L03
| |--+--Loberoschema bimaculata L03
| | `--Stengita L03
| `--+--Atomarops L03
| | |--A. curvitibialis L03
| | `--A. lewisi L03
| `--Toramus L03
| |--‘Tomarops’ bimaculatus L03
| |--T. hirtellus L03
| `--‘Tomarops’ longitarsis L03
`--Cryptophilini [Xenoscelinini] L03
| i. s.: Chinophagus Ljubarsky 1997 L03
|--Brachypterosa peckorum L03
`--+--+--Crowsonguptus mexicanus L03
| `--Cryptophilus Reitter 1874 [incl. Tomarops Grouvelle 1903] L03
| |--*C. integer (Heer 1841) [=Cryptophagus integer] L03
| |--C. allotrius [=Cryptophagops allotrius] L03
| |--C. leonensis [=Cryptophagops leonensis] L03
| |--C. mnionomoides [=Cryptophagops mniomonoides] L03
| `--‘*Tomarops’ punctatus Grouvelle 1903 L03
`--+--Cathartocryptus Sharp 1886 [incl. Xenoscelinus Grouvelle 1910] L03
| |--C. maculosus (Broun 1881) (see below for synonymy) L03
| `--‘*Xenoscelinus’ malaicus Grouvelle 1910 L03
`--Loberopsyllus L03
|--L. explanatus Leschen & Ashe 1999 B14
|--L. halffteri B14
|--L. oculatus L03
`--L. traubi B14
Cathartocryptus maculosus (Broun 1881) [=Paramecosoma maculosa, Xenoscelinus maculosus; incl. *C. obscurus Sharp 1886] L03
*Type species of generic name indicated
REFERENCES
[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).
[L03] Leschen, R. A. B. 2003. Erotylidae (Insecta: Coleoptera: Cucujoidea): phylogeny and review. Fauna of New Zealand 47: 1–108.
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