Scatopsoidea

Black compost fly Scatopse notata, copyright Miroslav Deml.

Belongs within: Bibionomorpha.

The Scatopsoidea are a clade of flies united by wings with M₁₊₂ fused to Rs for a distance, and M₃ and im lost; at least living species also have the larval head greatly reduced (Blagoderov et al. 2002). Members of the Scatopsidae have a single-segmented palpus and five to ten antennal flagellomeres (Amorim 2009) whereas Synneuridae have a four-segmented palpus and ten to fourteen flagellomeres (Peterson & Cook 1981).

<==Scatopsoidea BLM02
    |--Mesoscatopse RJ93 [Protoscatopsidae BLM02]
    |    `--M. rohdendorfi Kovalev in Kalugina & Kovalev 1985 RJ93
    |--Synneuridae [Canthyloscelidae, Hyperoscelididae] BLM02
    |    |--Exiliscelis californiensis PC81
    |    |--Prohyperoscelis jurassicus Kovalev in Kalugina & Kovalev 1985 RJ93
    |    `--Synneuron PC81
    |         |--S. annulipes PC81
    |         |--S. decipiens PC81
    |         `--S. silvestre PC81
    `--Scatopsidae A09
         |  i. s.: Pocolohostema Cook 1971 H02
         |         Parmaferia O98
         |         Aldrorardiella Enderlein 1936 H02
         |--Ectaetia [Ectaetiinae] A09
         |    `--E. clavipes C81
         |--Aspistinae A09
         |    |--Arthria analis A09, C81
         |    `--Aspistes A09
         |         |--A. berolinensis G20
         |         `--A. harti A09
         |--Psectrosciarinae A09
         |    |--Anapausis soluta A09
         |    `--Psectrosciara A09
         |         |--P. californica A09
         |         `--P. serrata A09
         `--Scatopsinae A09
              |  i. s.: Sinoscatopse Hong 2002 H02
              |           `--*S. eocenica Hong 2002 H02
              |--Colobostematini A09
              |    |--Colobostema variatum A09
              |    |--Borneoscatopse A09
              |    |--Holoplagia guamensis A09
              |    |--Lumpuria A09
              |    |--Cookella albitarsis A09
              |    `--Ferneiella A09
              |--Scatopsini A09
              |    |--Reichertella Enderlein 1912 A09, H02
              |    |--Apiloscatopse uncinata A09
              |    |--Pharsoreichertella A09
              |    `--Scatopse Greoffroy 1762 H02
              |         |--S. fuscipes CM91
              |         |--S. nigra L02
              |         `--S. notata [=Bibio (Scatopse) notatus] G20
              |--Rhegmoclematini A09
              |    |--Diamphidicus [Diamphidicina] A09
              |    |--Holoclema [Holoclematina] A09
              |    |--Rhegmoclematina A09
              |    |    |--Parmaferia A09
              |    |    |--Thripomorpha truncatum [=Rhegmoclema truncatum] A09
              |    |    `--Rhegmoclema Enderlein 1936 H02
              |    `--Parascatopsina A09
              |         |--Parascatopse wirthi A09
              |         |--Austroclemina A09
              |         |--Neorhegmoclemina Cook 1955 A09, H02
              |         |    `--N. asymmetrica A09
              |         `--Rhegmoclemina Enderlein 1936 A09, H02
              |              `--R. bimaculata A09
              `--Swammerdamellini A09
                   |--Akorhexoza cactivora A09
                   |--Coboldia fuscipes A09
                   |--Pararhexosa A09
                   |--Hawomersleya A09
                   |--Abrhexoza A09
                   |--Cooka A09
                   |--Brahemyia A09
                   |--Swammerdamella Enderlein 1912 H02
                   |    `--S. obtusa A09
                   |--Quateiella A09
                   |    |--Q. inexpectata A09
                   |    `--Q. minuscula A09
                   `--Rhexoza A09
                        |--R. incisa A09
                        `--R. similis A09

*Type species of generic name indicated

REFERENCES

[A09] Amorim, D. de S. 2009. Scatopsidae (minute black scavenger flies). In: Brown, B. V., A. Borkent, J. M. Cumming, D. M. Wood, N. E. Woodley & M. A. Zumbado (eds) Manual of Central American Diptera vol. 1 pp. 347–355. NRC Research Press: Ottawa.

[BLM02] Blagoderov, V. A., E. D. Lukashevich & M. B. Mostovski. 2002. Order Diptera Linné, 1758. The true flies (=Muscida Laicharting, 1781). In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 227–240. Kluwer Academic Publishers: Dordrecht.

[CM91] Colless, D. H., & D. K. McAlpine. 1991. Diptera (flies). In: CSIRO. The Insects of Australia: A textbook for students and research workers 2^nd ed. vol. 2 pp. 717–786. Melbourne University Press: Carlton (Victoria).

[C81] Cook, E. F. 1981. Scatopsidae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 313–319. Research Branch, Agriculture Canada.

[G20] Goldfuss, G. A. 1820. Handbuch der Naturgeschichte vol. 3. Handbuch der Zoologie pt 1. Johann Leonhard Schrag: Nürnberg.

[H02] Hong Y. 2002. Amber Insect of China. Beijing Scientific and Technological Publishing House.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[O98] Oosterbroek, P. 1998. The Families of Diptera of the Malay Archipelago. Brill: Leiden.

[PC81] Peterson, B. V., & E. F. Cook. 1981. Synneuridae. In: McAlpine, J. F., B. V. Peterson, G. E. Shewell, H. J. Teskey, J. R. Vockeroth & D. S. Wood (eds) Manual of Nearctic Diptera vol. 1 pp. 321–324. Research Branch, Agriculture Canada.

[RJ93] Ross, A. J., & E. A. Jarzembowski. 1993. Arthropoda (Hexapoda; Insecta). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 363–426. Chapman & Hall: London.

Last updated: 4 February 2021.