Kirkbyoidea

Amphissites alticostatus, (clockwise from top left) left, right, right interior and left interior valves, from Hoare & Merrill (2004).

Belongs within: Ostracoda.
Contains: Kirkbya, Polytylites.

The Kirkbyoidea are a group of reticulate, straight-backed ostracods known from the Devonian to the Middle Permian. In many examples, including members of the families Kirkbyidae, Amphissitidae and Arcyzonidae, the carapace valves are marked by a large subcentral pit (Benson et al. 1961).

Characters (from Benson et al. 1961, as Kirkbyacea): Reticulate, straight-backed, with or without lobes, nodes and carinae; ridge-and-groove hingement, with or without terminal dentition; valves subequal, overlap slight; free margin of one valve rabbeted to receive opposing valve; one or more marginal rims, dimorphism unknown.

<==Kirkbyoidea [Kirkbyacea, Kirkbyocopina]
    |--Cardiniferella Sohn 1953 [Cardiniferellidae] BB61
    |    |--*C. bowsheri Sohn 1953 BB61
    |    `--C. ringwoodensis (Harris & Jobe 1956) WSB93
    |--Kellettinidae [Kellettininae] HM04
    |    |--Semipetasus Sohn 1954 BB61
    |    |    `--*S. signatus Sohn 1954 BB61
    |    |--Kindlella Sohn 1954 HM04
    |    |    |--*K. fissiloba Sohn 1954 BB61
    |    |    `--K. proscillata Hoare & Merrill 2004 HM04
    |    `--Kellettina Swartz 1936 BB61
    |         |--*K. robusta (Kellett 1933) [=Ulrichia robusta] BB61
    |         `--K. vidriensis Hamilton 1942 WSB93
    |--Arcyzonidae ZL87
    |    |--Amphizona Kesling & Copeland 1954 ZL87
    |    |    `--*A. asceta Kesling & Copeland 1954 BB61
    |    |--Paegnium Kesling 1957 BB61
    |    |    `--*P. tanaum Kesling 1957 BB61
    |    |--Reticestus Kesling & Weiss 1953 BB61
    |    |    `--*R. acclivitatus Kesling & Weiss 1953 BB61
    |    |--Chironiptrum Kesling 1952 BB61
    |    |    |--*C. oiostathmicum Kesling 1952 BB61
    |    |    `--C. bilinearis Copeland 1977 WSB93
    |    `--Arcyzona Kesling 1952 BB61
    |         |--*A. diademata (Van Pelt 1933) [=Amphissites diadematus] BB61
    |         `--A. aperticarinata BB61
    |--Scrobiculidae [Placideidae] ZL87
    |    |--Scrobicula Posner 1951 BB61
    |    |    `--*S. scrobiculata (Jones, Kirkby & Brady 1884) [=Cytherella scrobiculata] BB61
    |    |--Bideirella Stover 1956 BB61
    |    |    `--*B. reticulata Stover 1956 BB61
    |    |--Amphissella Stover 1956 BB61
    |    |    `--*A. papillosa Stover 1956 BB61
    |    |--Doraclatum Stover 1956 BB61
    |    |    `--*D. compandium Stover 1956 BB61
    |    |--Placidea Schneider 1956 WSB93, BB61
    |    |    |--*P. lutkevichi (Spizharsky 1939) [=Amphissites lutkevichi] BB61
    |    |    `--P. trituberculata BB61
    |    `--Roundyella Bradfield 1935 HM04 [incl. Scaberina Bradfield 1935 BB61]
    |         |--*R. simplicissima (Knight 1928) BB61 [=Amphissites simplicissimus ZL87]
    |         |--R. bellatula Bradfield 1935 HM04
    |         |--R. nodomarginata (Bradfield 1935) HM04
    |         |--R. papilliformis (Wang 1978) CB86
    |         `--R. simplex (Girty 1910) HM04
    |--Kirkbyidae [Kirkbijidae] HM04
    |    |--Kirkbya HM04
    |    |--Villozona aspera (Polenova 1974) WSB93
    |    |--Carinaknightina carinata Sohn 1970 WSB93
    |    |--Coronakirkbya Sohn 1954 ZL87
    |    |    `--*C. fimbriata Sohn 1954 BB61
    |    |--Knightina Kellett 1933 [incl. Tenebrion Zanina 1956] BB61
    |    |    `--*K. allorismoides (Knight 1928) [=Amphissites allorismoides] BB61
    |    |--Editia Brayer 1952 M61, BB61
    |    |    `--*E. elegantis Brayer 1952 BB61
    |    |--Strepula Jones & Holl 1886 M61, BB61
    |    |    `--*S. concentrica Jones & Holl 1886 BB61
    |    `--Aurikirkbya Sohn 1950 HM04
    |         |--*A. wordensis (Hamilton 1942) [=Kirkbya wordensis] BB61
    |         |--A. kellettae (Harlton 1929) [=Kirkbya kellettae, Knightina kellettae] HM04
    |         |--A. knighti (Harlton 1929) HM04
    |         `--A. triserata Shaver in Thompson et al. 1959 HM04
    `--Amphissitidae HM04
         |--Polytylites HM04
         |--Vitissites comtei Becker 1981 WSB93
         |--Neoamphissites costatus Becker & Wang 1992 WSB93
         |--Kegelites Coryell & Booth 1933 HM04 (see below for synonymy)
         |    |--*K. spinosus (Coryell & Booth 1933) [=*Girtyites spinosus] BB61
         |    |--K. dattonensis (Harlton 1927) [=Amphissites dattonensis] ZL87
         |    |--K. harltoni (Cooper 1946) [=Ectodemites harltoni] HM04
         |    `--K. wapanuckensis (Harlton 1929) HM04
         |--Shleesha Sohn 1961 HM04
         |    |--S. houae (Chen 1958) [=Amphissites houae] CB86
         |    |--S. pinguis (Ulrich & Bassler 1906) [=Kirkbya pinguis] HM04
         |    |--S. pinguoides (Croneis & Gale 1939) HM04
         |    |--S. simplex (Roth 1929) HM04
         |    `--S. sullivanensis (Payne 1937) HM04 [=Shleesha sullivanensis ZL87, Ectodemites sullivanensis ZL87]
         `--Amphissites Girty 1910 HM04 (see below for synonymy)
              |  i. s.: A. centronotus (Ulrich & Bassler 1906) ZL87 (see below for synonymy)
              |         A. primus [=*Ectodemites primus] BB61
              |         A. robertsi Morey 1935 ZL87
              `--A. (Amphissites) HM04
                   |--*A. (A.) rugosus Girty 1910 BB61
                   |--A. (A.) alticostatus Bradfield 1935 HM05
                   |--A. (A.) insignis Croneis & Thurman 1939 HM04
                   |--A. (A.) morrowanensis Hoare & Merrill 2004 HM04
                   `--A. (A.) reticulatus Geis 1932 HM04

Amphissites Girty 1910 HM04 [incl. Albanella Harris & Lalicker 1932 BB61, Binodella Bradfield 1935 BB61, Brillius Brayer 1952 BB61, Ectodemites Cooper 1941 BB61]

Amphissites centronotus (Ulrich & Bassler 1906) ZL87 [=Kirkbya centronota ZL87; incl. A. blumenstengeli Gründel 1962 S82, A. mosquensis Posner 1951 S82]

Kegelites Coryell & Booth 1933 HM04 [=Girtyites Coryell & Booth 1933 non Wedekind 1914 BB61; incl. Kirkbyites Johnson 1936 BB61]

*Type species of generic name indicated

REFERENCES

[BB61] Benson, R. H., J. M. Berdan, W. A. van den Bold, T. Hanai, I. Hessland, H. V. Howe, R. V. Kesling, S. A. Levinson, R. A. Reyment, R. C. Moore, H. W. Scott, R. H. Shaver, I. G. Sohn, L. E. Stover, F. M. Swain & P. C. Sylvester-Bradley. 1961. Systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q99–Q421. Geological Society of America and University of Kansas Press.

[CB86] Chen D.-Q. & Bao H. 1986. Lower Permian ostracodes from the Chihsia Formation of Jurong and Longtan, Jiangsu Province. Acta Micropalaeontologica Sinica 3 (2): 107–132.

[HM04] Hoare, R. D., & G. K. Merrill. 2004. A Pennsylvanian (Morrowan) ostracode fauna from Texas. Journal of Paleontology 78 (1): 185–204.

[M61] Moore, R. C. 1961. Supplement—USSR treatise on Ostracoda. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt Q. Arthropoda 3: Crustacea: Ostracoda pp. Q422–Q429. Geological Society of America and University of Kansas Press.

[S82] Shi C.-G. 1982. Some early Carboniferous ostracodes from Nylam, Xizang (Tibet). Acta Palaeontologica Sinica 21 (3): 309–314.

[WSB93] Whatley, R. C., D. J. Siveter & I. D. Boomer. 1993. Arthropoda (Crustacea: Ostracoda). In: Benton, M. J. (ed.) The Fossil Record 2 pp. 343–356. Chapman & Hall: London.

[ZL87] Zhang X.-J. & Liang X.-Y. 1987. Ostracoda from the Taiyuan Formation of Xingyang and Gong Xian districts, Henan. Acta Micropalaeontologica Sinica 4 (3): 293–312.

Last updated: 17 October 2020.