Belongs within: Rubiaceae.
Contains: Naucleeae.
The Cinchonoideae as recognised here is a clade primarily supported by molecular analyses. Members of the clade have imbricate aestivation, are mostly woody, and many contain complex indole alkaloids (Bremer et al. 1999).
<==Cinchonoideae [Antirrheoideae]
| i. s.: Hillia [Hillioideae] T00
| |--H. parasitica J87
| |--H. triflora BM00
| `--H. valeris BM00
| Deppea grandiflora BM00, BJ99
| Steenisia BM00
|--Cinchona RB01
| |--C. pubescens RB01
| `--C. succirubra B96
`--+--+--+--Naucleeae RB01
| | `--Hymenodictyoneae RB01
| | |--Paracorynanthe antakarana RB01
| | `--Hymenodictyon RB01
| | |--H. decaryi RB01
| | |--H. flaccidum RB01
| | |--H. floribundum RB01
| | `--H. parvifolium RB01
| `--+--Strumpfia maritima BJ99
| |--Catesbaea [Catesbaeeae] B96
| | |--C. ekmaniana J87
| | `--C. spinosa BJ99
| `--Chiococceae B96
| | i. s.: Hintonia latiflora B96
| |--+--Chiococca alba BJ99
| | `--Erithalis fruticosa BJ99
| `--Exostema RB01
| |--E. caribaeum BJ99
| |--E. lineatum RB01
| `--E. picardae J87
`--+--Rondeletieae B96
| |--Rogiera suffrutescens B96, BJ99
| `--Rondeletia RB01
| |--R. carnea J87
| |--R. christii [incl. R. selleana] J87
| |--R. domatiata J87
| |--R. formonia J87
| |--R. leucophylla RB01
| `--R. odorata BJ99
`--+--Hamelieae B96
| |--Hoffmannia B96
| | |--H. ghiesbreghtii B96
| | `--H. refulgens B96
| `--Hamelia B96
| |--H. cuprea BM00
| |--H. papillosa BM00
| |--H. patens J87
| `--H. ventricosa J87
`--Guettardeae B96
|--Antirhea B96
| |--A. lucida BJ99
| `--A. obtusifolia SWK87
`--Guettarda B96
|--G. ovalifolia J87
|--G. pungens J87
|--G. speciosa P88
|--G. torbeciana J87
`--G. uruquensis BJ99
*Type species of generic name indicated
REFERENCES
[B96] Bremer, B. 1996. Combined and separate analyses of morphological and molecular data in the plant family Rubiaceae. Cladistics 12: 21-40.
[BJ99] Bremer, B., R. K. Jansen, B. Oxelman, M. Backlund, H. Lantz & K.-J. Kim. 1999. More characters or more taxa for a robust phylogeny – case study from the coffee family (Rubiaceae). Systematic Biology 48 (3): 413-435.
[BM00] Bremer, B., & J.-F. Manen. 2000. Phylogeny and classification of the subfamily Rubioideae (Rubiaceae). Plant Systematics and Evolution 225: 43-72.
[J87] Judd, W. S. 1987. Floristic study of Morne La Visite and Pic Macaya National Parks, Haiti. Bulletin of the Florida State Museum – Biological Sciences 32 (1): 1-136.
[P88] Polunin, I. 1988. Plants and Flowers of Malaysia. Times Editions: Singapore.
[RB01] Razafimandimbison, S. G., & B. Bremer. 2001. Tribal delimitation of Naucleeae (Cinchonoideae, Rubiaceae): inference from molecular and morphological data. Systematics and Geography of Plants 71: 515-538.
[SWK87] Snyder, N. F. R., J. W. Wiley & C. B. Kepler. 1987. The Parrots of Luquillo: Natural history and conservation of the Puerto Rican parrot. Western Foundation of Vertebrate Zoology: Los Angeles.
[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: Dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441-647.
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