Stygnus

Stygnus sp., copyright Arthur Anker.

Belongs within: Stygnidae.

Stygnus is a genus of short-legged harvestmen found in tropical South America.

Characters (from Pinto-da-Rocha 1997): Dorsal scute slightly wider at groove III. Cephalothorax with one central spine. Eye mounds smooth. Area I divided by median groove; III with two spines. Free tergite III with two tubercles larger than others. Tarsal process present. Scopula usually absent (present in S. gertschi). Pedipalpal femur large; basal sockets at apex of tibia normal. Penis ventral plate with lateral margins parallel; three, rarely four, pairs of distal (wide and large) and three or four basal pairs of setae present; dorsal process present.

<==Stygnus Perty 1833 (see below for synonymy) K03
    |--*S. armatus Perty 1833 K03
    |--S. aggerum Sørensen 1932 [=Metastygnellus aggerum; incl. *Chelistygnus bellicosus Roewer 1943] K03
    |--S. brevispinis Pinto-da-Rocha 1997 K03
    |--S. ferrugineus (Perty 1832) DG09 [=Goniosoma ferrugineum K03, *Stygnellus ferrugineus K03]
    |--S. gertschi (Roewer 1963) [=*Satiastygnus gertschi] K03
    |--S. grasshoffi Pinto-da-Rocha 1997 K03
    |--S. heliae Pinto-da-Rocha 1997 K03
    |--S. klugi (Goodnight & Goodnight 1943) [=*Parajanellus klugi] K03
    |--S. lesserti (Roewer 1943) [=*Stygnellomma lesserti] K03
    |--S. luteus (Mello-Leitão 1931) (see below for synonymy) K03
    |--S. marthae Pinto-da-Rocha 1997 K03
    |--S. mediocris (Roewer 1931) [=*Lojanellus mediocris] K03
    |--S. multispinosus (Piza 1938) [=Metastygnellus multispinosus] K03
    |--S. pectinipes (Roewer 1943) [=*Stygnellops pectinipes] K03
    |--S. peruvianus (Roewer 1957) [=Lojanellus peruvianus] K03
    |--S. polyacanthus (Mello-Leitão 1923) (see below for synonymy) K03
    |--S. simonis (Sørensen 1932) [=*Henriksia simonis] K03
    |--S. simplex (Roewer 1913) [=*Parastygnellus simplex] K03
    |--S. tocantinensis Pinto-da-Rocha 1997 K03
    `--S. weyrauchi (Roewer 1963) [=Lojanellus weyrauchi] K03

Stygnus Perty 1833 [incl. Acrostygnus Roewer 1931, Chelistygnus Roewer 1943, Diplostygnus Mello-Leitão 1931, Henriksia Mello-Leitão 1933, Lojanellus Roewer 1931, Metastygnellus Mello-Leitão 1928, Parajanellus Goodnight & Goodnight 1943, Parastygnellus Roewer 1913, Satiastygnus Roewer 1963, Stygnellomma Roewer 1943, Stygnellops Roewer 1943, Stygnellus Roewer 1913, Stygnophareus Soares 1970] K03

Stygnus luteus (Mello-Leitão 1931) [=*Diplostygnus luteus; incl. D. leonardosi Mello-Leitão 1939, *Stygnophareus novaisi Soares 1970, D. tibialis Mello-Leitão 1941] K03

Stygnus polyacanthus (Mello-Leitão 1923) [=Stignellus (l. c.) polyacanthus, *Acrostygnus polyacanthus; incl. *Metastygnellus longimanus Mello-Leitão 1928, Metartygnellus (l. c.) longimanus, Metas. nasutus Mello-Leitão 1944] K03

*Type species of generic name indicated

REFERENCES

[DG09] DaSilva, M. B., & P. Gnaspini. 2009. A systematic revision of Goniosomatinae (Arachnida: Opiliones: Gonyleptidae), with a cladistic analysis and biogeographical notes. Invertebrate Systematics 23: 530–624.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

Pinto-da-Rocha, R. 1997. Systematic review of the Neotropical family Stygnidae (Opiliones, Laniatores, Gonyleptoidea). Arq. Zool., Sao Paulo 33 (4): 163–342.