Stygnopsidae

Stygnopsis robusta, from here.

Belongs within: Grassatores.

The Stygnopsidae are a group of harvestmen found in Mexico and Central America characterised by male genitalia with an internal capsule forming a follis on the ventral side in dorsal view of the pars distalis, with a few distal espiniform projections and with several setae on the pars distalis (Cruz-López & Francke 2013).

Stygnopsidae [Isaeinae, Stygnopsinae]
    |--Karos Goodnight & Goodnight 1944 SG11, K03 (see below for synonymy)
    |    |--*K. barbarikos Goodnight & Goodnight 1944 K03
    |    |--K. brignolii Šilhavý 1974 K03
    |    |--K. depressus Goodnight & Goodnight 1971 K03
    |    |--K. dybasi (Goodnight & Goodnight 1947) [=*Potosa dybasi] K03
    |    |--K. gratiosus Goodnight & Goodnight 1971 K03 K03
    |    |--K. parvus Goodnight & Goodnight 1971 K03
    |    |--K. projectus Goodnight & Goodnight 1971 K03
    |    |--K. rugosus Goodnight & Goodnight 1971 K03
    |    `--K. tuberculatus (Goodnight & Goodnight 1944) [=*Monterella tuberculata] K03
    `--+--Hoplobunus Banks 1900 SG11, K03 (see below for synonymy)
       |    |--*H. barretti Banks 1900 [=H. baretti (l. c.)] K03
       |    |--H. apoalensis Goodnight & Goodnight 1973 K03
       |    |--H. boneti (Goodnight & Goodnight 1942) [=*Serrobunus boneti] K03
       |    |--H. mexicanus (Roewer 1915) [=*Haehnelia mexicana] K03
       |    |--H. neglectus Kury & Cokendolpher 2000 (see below for synonymy) K03
       |    |--H. oaxacensis Goodnight & Goodnight 1973 K03
       |    |--H. planus Goodnight & Goodnight 1973 K03
       |    |--H. queretarius Šilhavý 1974 K03
       |    |--H. spinooculorum Goodnight & Goodnight 1973 K03
       |    `--H. zullinii Šilhavý 1977 K03
       `--Stygnopsis Sørensen 1902 SG11, K03
            |--*S. valida (Sørensen 1884) [=Stygnus validus, Stygnopsis validus] K03
            `--S. robusta (Goodnight & Goodnight 1971) [=Hoplobunus robustus] K03

Stygnopsidae incertae sedis:
  Mexotroglinus Šilhavý 1977 K03
    `--*M. sbordonii Šilhavý 1977 K03
  Tampiconus Roewer 1949 K03
    `--*T. philippii Roewer 1949 K03
  Troglostygnopsis Šilhavý 1974 [Troglostygnopsinae] K03
    |--*T. anophthalma Šilhavý 1974 K03
    `--T. inops (Goodnight & Goodnight 1971) [=Hoplobunus inops] K03
  Sbordonia Šilhavý 1977 K03
    |--*S. armigera Šilhavý 1977 K03
    `--S. parvula (Goodnight & Goodnight 1953) [=Paramitraceras parvulus] K03
  Paramitraceras Pickard-Cambridge 1905 (see below for synonymy) K03
    |--*P. granulatum Pickard-Cambridge 1905 K03, KA-Z11, K03 (see below for synonymy)
    |--P. femorale Goodnight & Goodnight 1953 KA-Z11, K03 [=P. femoralis K03]
    `--P. hispidulum Pickard-Cambridge 1905 KA-Z11, K03 [=*Panzosus hispidulus K03, Par. hispidulus K03]
  Chinquipellobunus Goodnight & Goodnight 1944 C04
    |--*C. osorioi Goodnight & Goodnight 1944 [=Hoplobunus osorioi] C04
    |--C. coahuilaensis Cokendolpher 2004 C04
    |--C. madlae (Goodnight & Goodnight 1967) [=Hoplobunus madlae] C04
    |--C. mexicanus (Goodnight & Goodnight 1971) [=Pellobunus mexicanus] C04
    `--C. russelli (Goodnight & Goodnight 1967) [=Hoplobunus russelli] C04
  Philora Goodnight & Goodnight 1954 C-LF13
    |--*P. tuxtlae Goodnight & Goodnight 1954 C-LF13
    `--P. quetzalzin Cruz-López & Francke 2013 C-LF13
  Chapulobunus Goodnight & Goodnight 1946 C-LF16
    |--*C. unispinosus Goodnight & Goodnight 1946 C-LF16 [=Karos unispinosus K03]
    |--C. asper Cruz-López & Francke 2016 C-LF16
    |--C. poblano C-LF16
    |--C. psilocybe Cruz-López & Francke 2016 C-LF16
    `--C. regiomontano Cruz-López & Francke 2016 C-LF16
  Montabunus Goodnight & Goodnight 1945 C-LF16
    `--*M. foliorum Goodnight & Goodnight 1945 [=Karos foliorum] K03

Hoplobunus Banks 1900 SG11, K03 [incl. Haehnelia Roewer 1915 K03, Isaeus Sørensen 1932 K03, Serrobunus Goodnight & Goodnight 1942 K03]

Hoplobunus neglectus Kury & Cokendolpher 2000 [incl. *Isaeus mexicanus Sørensen 1932, H. mexicanus non Haehnelia mexicana Roewer 1915] K03

Karos Goodnight & Goodnight 1944 SG11, K03 [incl. Monterella Goodnight & Goodnight 1944 K03, Potosa Goodnight & Goodnight 1947 K03]

Paramitraceras Pickard-Cambridge 1905 [=Paramitroceras (l. c.); incl. Panzosus Roewer 1949, Solola Roewer 1949] K03

*Paramitraceras granulatum Pickard-Cambridge 1905 K03, KA-Z11, K03 [=P. granulatus K03; incl. P. chichivaca Goodnight & Goodnight 1947 K03, *Solola robusta Roewer 1949 K03]

*Type species of generic name indicated

REFERENCES

[C04] Cokendolpher, J. C. 2004. Revalidation of the harvestman genus Chinquipellobunus (Opiliones: Stygnopsidae). Texas Memorial Museum, Speleological Monographs 6: 143–152.

[C-LF13] Cruz-López, J. A., & O. F. Francke. 2013. On the enigmatic genus Philora: familial assignment and taxonomic revision (Opiliones: Laniatores: Stygnopsidae). Journal of Arachnology 41: 291–305.

[C-LF16] Cruz-López, J. A., & O. F. Francke. 2016. Three new species of the Mexican harvestman genus Chapulobunus (Opiliones: Stygnopsidae). Journal of Arachnology 44: 65–75.

[K03] Kury, A. B. 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Ibérica de Aracnología, volumen especial monográfico 1: 1–337.

[KA-Z11] Kury, A. B., & M. A. Alonso-Zarazaga. 2011. Addenda and corrigenda to the "Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones)". Zootaxa 3034: 47–68.

[SG11] Sharma, P. P., & G. Giribet. 2011. The evolutionary and biogeographic history of the armoured harvestmen—Laniatores phylogeny based on ten molecular markers, with the description of two new families of Opiliones (Arachnida). Invertebrate Systematics 25: 106–142.