Brachyceridae

Aonychus sp., copyright Irene Richardson.


Belongs within: Coleoptera.
Contains: Erirhinini, Hydronomini.

The Brachyceridae are a group of weevils mostly associated with water or damp habitats. They are superficially similar in external appearance to members of the family Curculionidae but may be distinguished by the more plesiomorphic nature of their genitalia, with males having a dorsally mostly membranous aedeagus, and a tegmen about as long or longer than the aedeagus and its apodemes (Zimmerman 1993).

<==Brachyceridae [Erirhinidae] B14
    |  i. s.: Tanysphyrus Z93
    |         Brachybamus Z93
    |         Penestes Z93
    |--Raymondionyminae B14
    |    |--Raymondionymus perrisi B14
    |    |--Schizomicrus B14
    |    |--Bordoniola B14
    |    `--Alaocybites B14
    |         |--A. californicus Gilbert 1956 B14
    |         |--A. egorovi B14
    |         `--A. rothi B14
    |--Tadiinae Z93
    |    |--Tadius Pascoe 1885 [Tadiini] Z93
    |    |    |--*T. erirhinoides Pascoe 1885 (see below for synonymy) Z93
    |    |    |--T. barbatus Lea 1928 [=T. barbatulus (l. c.)] Z93
    |    |    `--T. laticollis Faust 1898 Z93
    |    `--Aonychus Schoenherr 1844 [Aonychini] Z93
    |         |--*A. hopei Boheman 1844 M86
    |         |--A. barbatus Lea 1927 Z93
    |         |--A. bicruciatus Lea 1905 [=A. hopei var. bicruciatus] Z93
    |         |--A. lineatus Pascoe 1870 M86
    |         |--A. lituratus Lea 1927 Z93
    |         |--A. luctuosus Pascoe 1872 [incl. A. picatus Lea 1927] Z93
    |         |--A. pachypus Lea 1905 Z93
    |         `--A. striatus Lea 1905 Z93
    |--Brachycerus [Brachycerinae] B14
    |    |--B. algirus L02
    |    |--B. monachus M94
    |    |--B. muricatus B14
    |    |--B. ornatus (Drury 1773) B14
    |    |--B. scutellaris Chevrolat 1846 E12
    |    |--B. semituberculatus Chevrolat 1846 E12
    |    `--B. tetanicus Chevrolat 1847 E12
    `--Erirhininae [Erirhinides] B14
         |  i. s.: Clisis modesta Pascoe 1875 M86
         |         Diethusa fervida Pascoe 1873 M86
         |         Eniopea amoena Pascoe 1873 M86
         |         Nedyleda semiusta Pascoe 1872 M86
         |         Oenochroma rubeta Pascoe 1872 M86
         |         Misophryce hispida Pascoe 1873 M86
         |         Hedyopsis selligera Pascoe 1873 M86
         |         Methone ornata Pascoe 1875 M86
         |         Paryzeta musiva Pascoe 1873 M86
         |         Olanaea nigricollis Pascoe 1873 M86
         |         Cyttalia griseipila Pascoe 1873 M86
         |         Glaucopela unicolor Pascoe 1874 M86
         |         Empira variegata Pascoe 1874 M86
         |         Trechia Pascoe 1873 M86
         |         Agestra M86
         |           |--A. rubiginea Pascoe 1875 M86
         |           `--A. suturalis Pascoe 1873 M86
         |         Lybaeba M86
         |           |--L. repanda Pascoe 1873 M86
         |           `--L. subfasciata Pascoe 1873 M86
         |         Nemestra M86
         |           |--N. incerta Pascoe 1872 M86
         |           `--N. vibrata Pascoe 1882 M86
         |         Gerynassa M86
         |           |--G. basalis Pascoe 1873 M86
         |           `--G. nodulosa Pascoe 1873 M86
         |         Xeda M86
         |           |--X. amplipennis Pascoe 1873 M86
         |           `--X. bilineata Pascoe 1873 M86
         |         Orpha M86
         |           |--O. flavicornis Pascoe 1870 M86
         |           `--O. persimilis Pascoe 1872 M86
         |         Phaeodica M86
         |           |--P. fulvicornis Pascoe 1874 M86
         |           `--P. scutellaris Pascoe 1874 M86
         |         Phrenozemia M86
         |           |--P. lunata Pascoe 1873 M86
         |           `--P. lyproides Pascoe 1872 M86
         |         Enide M86
         |           |--E. aestuans Pascoe 1873 M86
         |           |--E. porphyrea Pascoe 1873 M86
         |           `--E. saniosa Pascoe 1873 M86
         |         Brexius M86
         |           |--B. angusticollis Pasoce 1870 M86
         |           |--B. lineatus Pascoe 1873 M86
         |           `--B. murinus Pascoe 1870 M86
         |         Antyllis M86
         |           |--A. aurulenta Pascoe 1873 M86
         |           |--A. griseola Pascoe 1873 M86
         |           |--A. setosa Pascoe 1873 M86
         |           `--A. togata Pascoe 1875 M86
         |         Emplesis M86
         |           |--E. filirostris Pascoe 1873 M86
         |           |--E. lineigera Pascoe 1870 M86
         |           |--E. scolopax Pascoe 1870 M86
         |           |--E. simplex Pascoe 1870 M86
         |           `--E. storeoides Pascoe 1873 M86
         |         Meriphus M86
         |           |--M. coronatus Pascoe 1873 M86
         |           |--M. fullo Erichs. 1842 M86
         |           |--M. guttatus Pascoe 1870 M86
         |           |--M. longirostris Pascoe 1871 M86
         |           `--M. umbrinus Pascoe 1870 M86
         |         Cydmaea M86
         |           |--C. bimaculata Pascoe 1872 M86
         |           |--C. luctuosa Pascoe 1872 M86
         |           |--C. notaticollis Pascoe 1875 M86
         |           |--C. pusilla Pascoe 1872 M86
         |           |--C. selligera Pascoe 1875 M86
         |           `--C. viridula Pascoe 1875 M86
         |         Rhachiodes M86
         |           |--R. bicaudatus (Boisduval 1835) [=Myllorhinus bicaudatus; incl. R. spinicollis] M86
         |           |--R. conformis Chev. 1879 M86
         |           |--R. dentifer [=Pteroporus dentifer] M86
         |           |--R. granulifera Chev. 1879 M86
         |           |--R. multidentatus Chev. 1879 M86
         |           |--R. nigropunctatus Chev. 1879 M86
         |           |--R. posticus Chev. 1879 M86
         |           `--R. signaticollis Chev. 1879 M86
         |         Aopolocnemis M86
         |           |--A. alboguttatus Chev. 1879 M86
         |           |--A. dorsonotatus Chev. 1879 M86
         |           |--A. fulvus Chev. 1879 M86
         |           |--A. guttiger Pascoe 1882 M86
         |           |--A. lineata Pascoe 1872 M86
         |           |--A. phaleratus Erichs. 1842 M86
         |           |--A. rufipes M86
         |           |--A. semicinctus Chev. 1879 M86
         |           `--A. suturalis Pascoe 1882 M86
         |         Lissorhoptrus M94
         |         Misophrice B70
         |--Erirhinini Z93
         |--Hydronomini Z93
         `--Neochetina Hustache 1926 B14, Z93 [Stenopelmi, Stenopelmini B14]
              |--*N. bruchi Hustache 1926 Z93
              `--N. eichhorniae Warner 1970 Z93

*Tadius erirhinoides Pascoe 1885 [incl. T. australiae Lea 1928, T. erirhinoides lineatus Faust 1894, T. erirhinoides suspiciosus Faust 1894] Z93

*Type species of generic name indicated

REFERENCES

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l’Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[M86] Masters, G. 1886. Catalogue of the described Coleoptera of Australia. Part V. Proceedings of the Linnean Society of New South Wales, series 2, 1 (3): 585–686.

[M94] May, B. M. 1994. An introduction to the immature stages of Australian Curculionoidea. In: Zimmerman, E. C. Australian Weevils (Coleoptera: Curculionoidea) vol. 2. Brentidae, Eurhynchidae, Apionidae and a chapter on immature stages by Brenda May pp. 365–728. CSIRO Australia.

[Z93] Zimmerman, E. C. 1993. Australian Weevils (Coleoptera: Curculionoidea) vol. 3. Nanophyidae, Rhynchophoridae, Erirhinidae, Curculionidae: Amycterinae, literature consulted. CSIRO Australia.

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