Omma stanleyi, copyright David Maddison.

Belongs within: Coleoptera.

The Archostemata are a distinctive group of wood-boring beetles with the labrum fused to the head capsule in adults, and a distinctive wing-tucking mechanism where the wing tips are rolled rather than folded under the elytra (Grimaldi & Engel 2005).

See also: Eating Mum from the inside out.

<==Archostemata [Cupedoidea]
    |  i. s.: Jurodidae B14
    |           |--Jurodes ignoramus B14
    |           `--Sikhotealinia [Sikhotealiniidae] B14
    |                `--S. zhiltzovae Lafer 1996 B14
    |         Obrienia P02 [Obrieniidae GE05]
    |           `--O. kuscheli P02
    |--+--Micromalthus [Micromalthidae, Micromalthinae, Micromalthoidea] MW15
    |  |    `---M. debilis LeConte 1878 PN02
    |  `--Ommatidae [Ommatinae, Tetraphaleridae] MW15
    |       |--Notocupes Ponomarenko 1964 KL01
    |       |--Tetraphalerus LM87
    |       |    |--T. bruchi HR11
    |       |    `--T. verrucosus P02
    |       |--Omamima GE05
    |       |--Tetraphalerites GE05
    |       |--Omma Newman 1839 LM87
    |       |    |--*O. stanleyi Newman 1839 LM87
    |       |    |--O. mastersi Macleay 1871 LM87
    |       |    |--O. sagitta B14
    |       |    `--O. varians B70
    |       `--Crowsoniella [Crowsoniellidae] LM87
    |            `--C. relicta Pace 1975 B14
    `--Cupedidae MW15
         |  i. s.: Triadocupedinae P02
         |         Mesocupedinae LM87
         |--Priacma [Priacminae] MW15
         |    `--P. serrata (LeConte 1861) B14
         `--+--Prolixocupes lobiceps MW15, HR11
            `--Cupedinae LM87
                 |  i. s.: Rhipsidegma raffrayi (Fairmaire 1884) B14
                 |         Adinolepis Neboiss 1984 LM87
                 |           |--*A. eumana (Neboiss 1960) [=Cupes eumana] LM87
                 |           |--A. mathesonae (Neboiss 1960) [=Cupes mathesonae] LM87
                 |           |--A. scalena Neboiss 1984 LM87
                 |           `--A. youanga (Neboiss 1960) [=Cupes youanga] LM87
                 |--Tenomerga MW15
                 |    |--T. concolor LM87
                 |    `--T. mucida LM87
                 `--+--Distocupes Neboiss 1984 MW15, LM87
                    |    `--*D. varians (Lea 1902) [=Cupes varians] LM87
                    `--Cupes capitatus Fabricius 1801 MW15, B14

*Type species of generic name indicated


[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[B70] Britton, E. B. 1970. Coleoptera (beetles). In: CSIRO. The Insects of Australia: A textbook for students and research workers pp. 495–621. Melbourne University Press.

[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.

[HR11] Heraty, J., F. Ronquist, J. M. Carpenter, D. Hawks, S. Schulmeister, A. P. Dowling, D. Murray, J. Munro, W. C. Wheeler, N. Schiff & M. Sharkey. 2011. Evolution of the hymenopteran megaradiation. Molecular Phylogenetics and Evolution 60: 73–88.

[KL01] Krzeminski, W., & C. Lombardo. 2001. New fossil Ephemeroptera and Coleoptera from the Ladinian (Middle Triassic) of Canton Ticino (Switzerland). Rivista Italiana di Paleontologia e Stratigrafia 107 (1): 69–78.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[MW15] McKenna, D. D., A. L. Wild, K. Kanda, C. L. Bellamy, R. G. Beutel, M. S. Caterino, C. W. Farnum, D. C. Hawks, M. A. Ivie, M. L. Jameson, R. A. B. Leschen, A. E. Marvaldi, J. V. McHugh, A. F. Newton, J. A. Robertson, M. K. Thayer, M. F. Whiting, J. F. Lawrence, A. Ślipiński, D. R. Maddison & B. D. Farrell. 2015. The beetle tree of life reveals that Coleoptera survived end-Permian mass extinction to diversify during the Cretaceous terrestrial revolution. Systematic Entomology 40 (4): 835–880.

[PN02] Pollock, D. A., & B. B. Normark. 2002. The life cycle of Micromalthus debilis LeConte (1878) (Coleoptera: Archostemata: Micromalthidae): historical review and evolutionary perspective. Journal of Zoological Systematics and Evolutionary Research 40 (2): 105–112.

[P02] Ponomarenko, A. G. 2002. Superorder Scarabaeidea Laicharting, 1781. Order Coleoptera Linné, 1758. The beetles. In: Rasnitsyn, A. P., & D. L. J. Quicke (eds) History of Insects pp. 164–176. Kluwer Academic Publishers: Dordrecht.

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