Pholcinae

Pholcus phalangioides, copyright Olaf Leillinger.

Belongs within: Pholcidae.

The Pholcinae are a group of daddy-long-legs spiders united by the reduction of the tarsal comb hairs to a single ventral row, and the usual presence on the male chelicerae of a pair of proximal lateral apophyses (Huber 2011).

<==Pholcinae
    |  i. s.: Aetana Huber 2005 H11
    |         Wanniyala Huber & Benjamin 2005 H11
    |--+--Nyikoa Huber 2007 H11
    |  |    `--N. limbe Huber 2007 HF08
    |  `--Zatavua Huber 2003 H11
    |       |--Z. vohiparara Huber 2003 HF08
    |       `--Z. zanahary H11
    `--+--Paraspermophora Wunderlich 2004 H11
       |--Anansus Huber 2007 H11
       |--Savarna Huber 2005 H11
       |--Khorata Huber 2005 H11
       |    `--K. khammouan Huber 2005 HF08
       |--+--+--Paramicromerys Millot 1946 H11
       |  |  |    `--P. scharffi Huber 2003 HF08
       |  |  `--+--Buitinga Huber 2003 HF08, H11
       |  |     |    |--B. asax Huber 2003 HF08
       |  |     |    `--B. kadogo H11
       |  |     `--Spermophorides Wunderlich 1992 HF08, H11
       |  |          |--S. cuneata (Wunderlich 1987) HF08
       |  |          `--S. elevata (Simon 1873) HF08
       |  `--+--Metagonia Simon 1893 H11
       |     |    |--M. conica (Simon 1893) HF08
       |     |    |--M. mariguitarensis (González-Sponga 1998) T09
       |     |    `--M. reventazona Huber 1997 HF08
       |     `--Spermophora Hentz 1841 HF08, H11
       |          |--S. kerinci Huber 2005 HF08
       |          |--S. senoculata (Dugès 1836) H11
       |          `--S. usambara Huber 2003 HF08
       `--+--Quamtana Huber 2003 H11
          |    `--Q. bonamanzi Huber 2003 HF08
          |--Belisana Thorell 1898 H11
          |    |--B. airai Huber 2005 HF08
          |    |--B. fiji Huber 2005 HF08
          |    |--B. hormigai H11
          |    `--B. ketambe H11
          `--+--Micropholcus Deeleman-Reinhold & Prinsen 1987 H11
             |    `--M. fauroti (Simon 1887) H11
             |--Pehrforsskalia Deeleman-Reinhold & van Harten 2001 H11
             |    `--P. conopyga Deeleman-Reinhold & van Harten 2001 HF08
             `--+--+--Panjange Deeleman-Reinhold & Deeleman 1983 HF08, H11
                |  |    `--P. mirabilis Deeleman-Reinhold 1986 HF08
                |  `--Leptopholceae H11
                |       |--Micromerys Bradley 1877 H11
                |       |    `--M. daviesae Deeleman-Reinhold 1986 HF08
                |       `--Leptopholcus Simon 1893 H11
                |            |--L. debakkeri H11
                |            |--L. gracilis H11
                |            `--L. hispaniola Huber 2000 HF08
                `--+--Calapnita Simon 1892 HF08, H11
                   |    |--C. phyllicola Deeleman-Reinhold 1986 HF08
                   |    `--C. saluang H11
                   |--Ossinissa Dimitrov & Ribera 2005 HF08, H11
                   |    `--*O. justoi (Wunderlich 1991) [=Spermophorides justoi] F05
                   |--Uthina Simon 1893 HF08, H11
                   |    `--U. luzonica Simon 1893 HF08
                   `--Pholcus Walckenaer 1805 H11
                        |--P. barbarus Lucas 1846 E12
                        |--P. bourgini H11
                        |--P. caudatus V09
                        |--P. chattoni H11
                        |--P. crypticolens Bösenberg & Strand 1906 RA08
                        |--P. kwamgumi H11
                        |--P. lamperti H11
                        |--P. manueli Gertsch 1937 [incl. P. affinis Schenkel 1953] RA08
                        |--P. opilionoides (Schrank 1781) HF08
                        |--P. phalangioides (Fuesslin 1775) H11
                        |--P. quadripunctatus Lucas 1846 E12
                        |--P. quinquenotatus H11
                        |--P. sphaeroides B96
                        `--P. zichyi Kulczyński 1901 K01

*Type species of generic name indicated

REFERENCES

[B96] Brunet, B. 1996. Spiderwatch: A Guide to Australian Spiders. Reed New Holland: Sydney.

[E12] Evenhuis, N. L. 2012. Publication and dating of the Exploration Scientifique de l'Algérie: Histoire Naturelle des Animaux Articulés (1846–1849) by Pierre Hippolyte Lucas. Zootaxa 3448: 1–61.

[F05] Fernández, J. 2005. Noticia de nuevos táxones para la ciencia en el ámbito Íbero-Balear y Macaronésico. Nuevos táxones animales descritos en la península Ibérica y Macaronesia desde 1994 (IX). Graellsia 61 (2): 261–282.

[H11] Huber, B. A. 2011. Phylogeny and classification of Pholcidae (Araneae): an update. Journal of Arachnology 39 (2): 211–222.

[HF08] Huber, B. A., & N. Fleckenstein. 2008. Comb-hairs on the fourth tarsi in pholcid spiders. Journal of Arachnology 36 (2): 232–240.

[K01] Kulczyński, V. 1901. Arachnoideák [Arachnoidea]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 311–369. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[RA08] Ramalho, M. O., D. Araujo, M. C. Schneider, A. D. Brescovit & D. M. Cella. 2008. Mesabolivar brasiliensis (Moenkhaus 1898) and Mesabolivar cyaneotaeniatus (Keyserling 1891) (Araneomorphae, Pholcidae): close relationship reinforced by cytogenetic analysis. Journal of Arachnology 36 (2): 453–456.

[T09] Taylor, C. K. 2009. Australiscutum, a new genus of Monoscutidae (Arachnida: Opiliones) from eastern Australia, with the first record of asymmetrical chelicerae in Opiliones. Insect Systematics and Evolution 40: 319–332.

[V09] Verdcourt, B. (ed.) 2009. Additions to the Wild Fauna and Flora of the Royal Botanic Gardens, Kew. XXVI. Miscellaneous records. Kew Bulletin 64 (1): 183–194.