Forest caterpillar hunter Calosoma sycophanta, copyright Thomas Huntke.

Belongs within: Carabinae.

The Carabini are a group of ground beetles with open procoxal and metacoxal cavities, and contiguous metacoxae.

Carabini B90
    |--Calosoma Weber 1801 B90, LM87
    |    |  i. s.: C. australe Hope 1845 LM87
    |    |         C. cyaneoventre Mandl 1955 LM87
    |    |         C. frigidum B14
    |    |         C. inquisitor L02
    |    |         C. scrutator B14
    |    |--*C. (Calosoma) sycophanta (Linnaeus 1758) [=Carabus sycophanta] LM87
    |    `--C. (Australodrepa Lapouge 1929) LM87
    |         |--C. (A.) oceanicum Perroud 1864 (see below for synonymy) LM87
    |         `--C. (A.) schayeri Erichson 1842 [incl. C. curtisi Hope 1845, C. grandipenne Castelnau 1867] LM87
    `--Carabus Linnaeus 1758 [Carabina] L58
         |  i. s.: C. americanus Linnaeus 1758 L58
         |         C. auratus L02
         |         C. coriaceus Linnaeus 1758 L58
         |         C. crepitans Linnaeus 1758 L58
         |         C. cruxminor Linnaeus 1758 L58
         |         C. ferrugineus Linnaeus 1758 L58
         |         C. glabratus L91
         |         C. granulatus Linnaeus 1758 L58 [incl. C. granulatus var. solskyanus C01]
         |         C. henningi [incl. C. henningi var. roeschkei Csiki 1901, C. henningi var. sajanicus Csiki 1901] C01
         |         C. hortensis Linnaeus 1758 L58
         |         C. inquisitor Linnaeus 1758 L58
         |         C. intricatus Linnaeus 1761 VPB11
         |         C. krueberi C01
         |         C. latus Linnaeus 1758 L58
         |         C. leucophthalmus Linnaeus 1758 L58
         |         C. lividus Linnaeus 1758 L58
         |         C. mannerheimi C01
         |         C. maurus C01
         |         C. melanocephalus Linnaeus 1758 L58
         |         C. menetriesi C01
         |         C. multipunctatus Linnaeus 1758 L58
         |         C. nemoralis E04
         |         C. nitens Linnaeus 1758 L58 [incl. C. nitens var. fennicus C01]
         |         C. piceus Linnaeus 1758 L58
         |         C. regalis [incl. C. regalis var. cyanicollis, C. regalis var. pasianix] C01
         |         C. sibiricus C01
         |         C. smaragdinus [incl. C. smaragdinus var. mandshuricus] C01
         |         C. spinipes Linnaeus 1758 L58
         |         C. vaporariorum Linnaeus 1758 L58
         |         C. violaceus Linnaeus 1758 L58
         `--C. (Ohomopterus) SS06
              |--C. (O.) dehaanii SS06
              |--C. (O.) iwawakianus SS06
              |--C. (O.) uenoi (Ishikawa 1960) SS06, I92
              |--C. (O.) yaconinus SS06
              `--C. (O.) yamato SS06

Calosoma (Australodrepa) oceanicum Perroud 1864 [incl. C. (*A.) timorense Chaudoir 1869, C. walkeri Waterhouse 1898] LM87

*Type species of generic name indicated


[B90] Bell, R. T. 1990. Insecta: Coleoptera Carabidae adults and larvae. In: Dindal, D. L. (ed.) Soil Biology Guide pp. 1053–1092. John Wiley & Sones: New York.

[B14] Bouchard, P. (ed.) 2014. The Book of Beetles: A lifesize guide to six hundred of nature's gems. Ivy Press: Lewes (United Kingdom).

[C01] Csiki, E. 1901. Bogarak [Coleopteren]. In: Horváth, G. (ed.) Zichy Jenő Gróf Harmadik Ázsiai Utazása [Dritte Asiatische Forschungsreise des Grafen Eugen Zichy] vol. 2. Zichy Jenő Gróf Harmadik Ázsiai Utazásának Állattani Eredményei [Zoologische Ergebnisse der Dritten Asiatischen Forschungsreise des Grafen Eugen Zichy] pp. 75–120. Victor Hornyánszky: Budapest, and Karl W. Hierseman: Leipzig.

[E04] Emberson, R. M. 2004. A European ground beetle, Harpalus tardus (Panzer) (Coleoptera: Carabidae), in Canterbury, and the predisposition of European carabids to establish in exotic temperate locations. New Zealand Entomologist 27: 39–42.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[L02] Latreille, P. A. 1802. Histoire Naturelle, générale et particulière des crustacés et des insectes vol. 3. Familles naturelles des genres. F. Dufart: Paris.

[LM87] Lawrence, J. F., B. P. Moore, J. E. Pyke & T. A. Weir. 1987. Zoological Catalogue of Australia vol. 4. Coleoptera: Archostemata, Myxophaga and Adephaga. Australian Government Publishing Service: Canberra.

[L58] Linnaeus, C. 1758. Systema Naturae per Regna Tria Naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata. Laurentii Salvii: Holmiae.

[L91] Lundqvist, L. 1991. Rearing deutonymphs of Iphidosoma fimetarium (J. Müller), a mesostigmatic mite associated with carabid beetles. In: Schuster, R., & P. W. Murphy (eds) The Acari: Reproduction, development and life-history strategies pp. 447–452. Chapman & Hall: London.

[SS06] Sota, T., & M. Sasabe. 2006. Utility of nuclear allele networks for the analysis of closely related species in the genus Carabus, subgenus Ohomopterus. Systematic Biology 55 (2): 329–344.

[VPB11] Varet, M., J. Pétillon & F. Burel. 2011. Comparative responses of spider and carabid beetle assemblages along an urban-rural boundary gradient. Journal of Arachnology 39 (2): 236–243.

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