Emarginula fissura, copyright Alboran Shells.

Belongs within: Fissurellidae.

The Emarginulini, slit limpets, are a group of limpet-like gastropods with a more-or-less developed longitudinal slit running back from the front of the shell (sometimes only present ventrally).

See also: Marginal limpets.

    |--Clypidina Gray 1847 [Clypidinidae] BR05
    |    `--*C. notata (Linné 1767) [=Patella notata] KC60
    |--Rimula Defrance 1827 BR05 [=Rimularia Waldheim 1834 KC60; Rimulacea, Rimulidae]
    |    |--*R. blainvilli Defrance 1827 [=*Rimularia blainvilli] KC60
    |    |--R. exquisita Adams 1853 H09
    |    `--R. mexicana Berry 1969 PP78
    |--Zeidora Adams 1860 BR05 (see below for synonymy)
    |    |  i. s.: Z. lodderae Tate & May 1900 H09
    |    |         Z. maoria Powell 1937 P61
    |    |--*Z. (Zeidora) calceolina Adams 1860 KC60
    |    `--Z. (Nesta Adams 1870) KC60
    |         `--Z. (*N.) candida (Adams 1870) [=*Nesta candida] KC60
    |--Montfortula Iredale 1915 [incl. Plagiorhytis Fischer 1885 non Chaudoir 1848] KC60
    |    |--*M. rugosa (Quoy & Gaimard 1834) KC60 (see below for synonymy)
    |    |--M. chathamensis Finlay 1928 P61
    |    |--M. conoidea (Reeve 1842) [=Hemitoma conoidea] F27
    |    |--M. kaawaensis (Bartrum 1919) [=Tugalia kaawaensis] F27
    |    |--M. lyallensis Mestayer 1928 P61
    |    |--M. pulchra P95
    |    `--M. variegata Adams 1852 W78
    |--Hemitoma Swainson 1840 BR05 (see below for synonymy)
    |    |  i. s.: H. bella (Gabb 1865) [=Emarginula bella] O27
    |    |         ‘Emarginula’ clathrata Ad. & Reeve 1850 [=Subemarginula clathrata] H09
    |    |         ‘Emarginula’ cumingii Sowerby 1863 [=Subemarginula cumingii] H09
    |    |         H. golischae Dall 1916 O27
    |    |         H. yatesii Dall 1901 O27
    |    |--H. (Hemitoma) octoradiata (Gmelin 1791) KC60 (see below for synonymy)
    |    |--H. (Montfortia Récluz 1843) KC60
    |    |    `--H. (*M.) australis (Quoy & Gaimard 1834) [=Emarginula australis] KC60
    |    `--H. (Montfortista Iredale 1929) KC60
    |         `--H. (*M.) excentrica (Iredale 1929) [=*Montfortista excentrica] KC60
    `--Emarginula Lamarck 1801 BR05 [=Emarginulus Montfort 1810 KC60, Imarginula Gray 1821 KC60]
         |  i. s.: E. convexa Hedley 1907 H09
         |         E. delicatissima F27
         |         E. dennanti F27
         |         E. dilecta Adams 1852 H09
         |         E. elongata M62
         |         E. foveolata NO04
         |           |--E. f. foveolata NO04
         |           `--E. f. fujitai NO04
         |         E. incisura Adams 1852 H09
         |         E. micans Adams 1852 H09
         |         E. ostheimerae Abbott 1958 B59
         |         E. patula Cotton 1930 MG-H11
         |         E. striatula (Quoy & Gaimard 1834) [incl. E. valentior] P61
         |         E. variegata Adams 1852 H09
         |         E. wannonensis F27
         |--E. (Emarginula) KC60
         |    |--*E. (E.) fissura [=Patella fissura] P61
         |    `--E. (E.) conica Lamarck 1801 KC60
         |--E. (Altomarginula Haber 1932) KC60
         |    `--E. (*A.) desnoyersi Eudes-Deslongchamps 1842 KC60
         |--E. (Arginula Palmer 1937) KC60
         |    `--E. (*A.) arata Conrad 1933 KC60
         |--E. (Entomella Cossmann 1888) KC60
         |    `--E. (*E.) clypeata Lamarck 1803 KC60
         |--E. (Semperia Crosse 1867) KC60
         |    `--E. (*S.) paivana (Crosse 1867) [=*Semperia paivana] KC60
         |--E. (Subzeidora Iredale 1924) KC60
         |    `--E. (*S.) connectens Thiele 1924 KC60
         `--E. (Tauschia Haber 1932) KC60
              `--E. (*T.) orthogonia Tausch 1890 KC60

Hemitoma Swainson 1840 BR05 [=Subemarginula Gray 1847 KC60; incl. Siphonella Issel 1869 non Hagenow 1851; Hemitomidae, Hemitominae]

Hemitoma (Hemitoma) octoradiata (Gmelin 1791) KC60 [=Patella octoradiata KC60, Emarginula (Hemitoma) octoradiata O27; incl. P. tricostata Humphrey ex Sowerby 1823 non Gmelin 1791 KC60, *Hemitoma tricostata KC60, *Subemarginula tricostata KC60]

*Montfortula rugosa (Quoy & Gaimard 1834) KC60 [=Emarginula rugosa F27, Clypidina (*Montfortula) rugosa KC60, Hemitoma rugosa F27, Subemarginula rugosa H09]

Zeidora Adams 1860 BR05 [=Zidora Fischer 1885 BR05; incl. Crepiemarginula Seguenza 1880 KC60, Legrandia Beddome 1883 non Hanley 1872 KC60; Zeidoridae, Zidoridae]

*Type species of generic name indicated


[B59] Baker, B. B. 1959. Publications received: The marine mollusks of Grand Cayman Island, British West Indies. By R. Tucker Abbott. Nautilus 72 (4): 147–148.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[M62] Monniot, F. 1962. Recherches sur les graviers a Amphioxus de la région de Banyuls-sur-Mer. Vie et Milieu 13: 231–322.

[NO04] Nakano, T., & T. Ozawa. 2004. Phylogeny and historical biogeography of limpets of the order Patellogastropoda based on mitochondrial DNA sequences. Journal of Molluscan Studies 70: 31–41.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II pt. III. Stanford University Press: Stanford University (California).

[P95] Ponder, W. F. 1995. Book review: Australian Marine Shells. Prosobranch Gastropods. Parts 1 and 2. Molluscan Research 16: 97–102.

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[W78] Wells, F. E. 1978. Zonation of marine gastropods on a rocky intertidal shore in the Admiralty Gulf, Western Australia, with emphasis on the genus Nerita. Veliger 20 (3): 279–287.

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