Dexiarchia

Tethys leporina, from here.


Belongs within: Opisthobranchia.
Contains: Aeolidida.

The Dexiarchia are a clade of nudibranchs characterised by a right-laterally located anus and usually a more or less branched digestive gland. The basal genus Doridoxa retains an unbranched digestive gland similar to that found in other nudibranchs.

<==Dexiarchia [Actenidiacea]
    |--Doridoxa Bergh 1899 [Doridoxida, Doridoxidae, Doridoxoidea, Pseudoeuctenidiacea] BR05
    `--Cladobranchia [Abranchia, Cladohepatica] BR05
         |  i. s.: Charcotiidae BR05
         |           |--Charcotia Vayssière 1906 BR05
         |           `--Leminda Griffiths 1985 [Lemindidae] BR05
         |         Goniaeolis Sars 1861 [Goniaeolididae] BR05
         |         Hero Lovén in Alder & Hancock 1855 [Heroacea, Heroidae, Heroinae] BR05
         |         Pinufius Marcus & Marcus 1960 [Pinufiidae] BR05
         |--Aeolidida BR05
         |--Arminina [Euarminoidea] H01
         |    |--Arminoidea [Arminacea, Euarminida] BR05
         |    |    |--Doridomorphidae [Doridoeididae] BR05
         |    |    |    |--Doridoeides Eliot & Evans 1908 BR05
         |    |    |    `--Doridomorpha Eliot 1903 BR05
         |    |    `--Arminidae [Euarminacea] BR05
         |    |         |--Pleuroleura Bergh 1874 [Pleuroleuridae] BR05
         |    |         |--Heterodoris Verrill & Emerton 1882 [Heterodoridae, Heterodorididae] BR05
         |    |         |--Dermatobranchus van Hasselt 1824 [Dermatobranchidae, Dermatobranchinae] BR05
         |    |         |--Atthila Bergh 1899 [Atthilidae] BR05
         |    |         |--Histiomena convolvula (Lance 1962) PP78
         |    |         `--Armina Rafinesque 1814 (see below for synonymy) BR05
         |    |              `--A. californica (Cooper 1862) PP78
         |    `--Metarminoidea BR05
         |         |--Dirona MacFarland in Eliot 1905 H01, BR05 [Dironidae]
         |         |--Madrella Alder & Hancock 1864 [Madrellidae] BR05
         |         `--Proctonotidae (see below for synonymy) BR05
         |              |--Zephyrina Quatrefages 1844 BR05
         |              |--Proctonotus Alder 1844 [Proctonotinae] BR05
         |              |--‘Janus’ Verany 1844 non Stephens 1835 [Janina] BR05
         |              |--‘Venilia’ Alder & Hancock 1844 (see below for synonymy) BR05
         |              |--Antiopella Hoyle 1902 [=Antiopa Alder & Hancock 1848 non Meigen 1800] BR05
         |              |    `--A. fusca (O’Donoghue 1924) H01
         |              `--Janolus Bergh 1884 BR05
         |                   |--*J. australis P61
         |                   `--J. novozelandica (Eliot 1907) P61 [=Antiopella novozealandica F27]
         `--Dendronotina H01
              |  i. s.: Dotidae [Dotonidae] H01
              |           |--Idulia Leach in Gray 1852 [Iduliidae] BR05
              |           `--Doto Oken 1815 BR05
              |                |--D. coronata (Gmelin 1791) H01
              |                `--D. pinnatifida (Montagu 1804) H01
              `--Tritonioidea [Dendronotida, Dicerata, Duvauceliidae, Seribranchia] BR05
                   |--Hancockia Gosse 1877 [Hancockidae, Hancockiidae] BR05
                   |--Scyllaea Linnaeus 1758 [Scyllaeidae] BR05
                   |--Bornella Gray 1850 [Bornellidae] BR05
                   |    `--B. digitata Adams & Reeve 1850 H09
                   |--Dendronotus Alder & Hancock 1845 [Dendronotacea, Dendronotidae, Dendronotinae] BR05
                   |    |--D. arborescens H04
                   |    `--D. iris Cooper 1863 H01
                   |--Lomanotus Verany 1844 [Lomanotidae] BR05
                   |    `--L. genei (Vérany 1846) H01
                   |--Aranucidae BR05
                   |    |--Aranucus Odhner 1936 [Aranucidae] BR05
                   |    `--Marianina Pruvot-Fol 1931 [Marianinidae] BR05
                   |--Phylliroidae [Phylliroidae, Phyllirrhoea, Psilosomata] BR05
                   |    |--Nectophyllirhoe Hoffmann 1922 [Nectophyllirhoidae] BR05
                   |    |--‘Dactylopus’ Bonnevie 1921 nec Gill 1859 nec Claus 1862 [Dactylopodidae] BR05
                   |    |--Phylliroe Péron & Lesueur 1810 [=Phyllirhoe (l. c.)] BR05
                   |    |    `--*P. bucephala (Peron & Lesueur 1810) P61
                   |    `--Pellibranchus Ralph 1944 P61
                   |         `--*P. cinnabareus Ralph 1944 P61
                   |--Tritoniidae [Duvauceliidae, Tritoniana, Tritoniomorpha, Zonabranchiatae] BR05
                   |    |--Duvaucelia Risso 1826 BR05
                   |    |--Marionia MG-H11
                   |    |--Necromantes Gistl 1848 P61 (see below for synonymy)
                   |    |    |--N. flemingi Powell 1937 P61
                   |    |    `--N. incerta (Bergh 1904) P61 [=Tritonia incerta F27, Sphaerostoma incerta F27]
                   |    `--Tritonia Cuvier 1797 [Tritoniinae] BR05
                   |         |--T. diomedia KC11
                   |         `--T. hombergii H04
                   `--Tethydidae [Tethyadae, Tethydia, Tethymelibidae] BR05
                        |--‘Fimbria’ Bohadsch ex O’Donoghue 1926 non Mühlfeld 1811 [Fimbriidae] BR05
                        |--Chioraera Gould 1852 [Chioraeridae] BR05
                        |--Melibe Rang 1829 [=Meliboea (l. c.); Melibidae, Melibinae, Meliboeadae] BR05
                        |    |--M. bucephala Bergh 1902 BW09
                        |    |--M. fimbriata Alder & Hancock 1864 BD86
                        |    `--M. leonina (Gould 1852) PP78
                        `--Tethys Linnaeus 1767 BR05 [incl. Phoenicurus Rudolphi 1819 non Forster 1817 D56; Tethydinae]
                             |--*T. leporina O27
                             |--T. sparsinotata (Smith 1884) [=Aplysia sparsinotata] H09
                             `--T. tigrina (Rang. 1828) [=Aplysia tigrina] H09

Armina Rafinesque 1814 [incl. Diphyllidia Blainville 1819, Pleurophyllidia Meckel 1823; Armininae, Diphyllidiae, Diphyllidiidae, Pleurophyllidiidae] BR05

Proctonotidae [Antiopellidae, Antiopidae, Antiopinae, Janolidae, Pachygnatha, Zephyrinidae] BR05

Necromantes Gistl 1848 P61 [incl. Sphaerostoma Macgillivray 1843 non Rudolphi 1809 P61, BR05; Sphaerostomatidae, Sphaerostomidae]

‘Venilia’ Alder & Hancock 1844 nec Rafinesque 1815 nec Morton 1833 [Veniliinae, Venilinae] BR05

*Type species of generic name indicated

REFERENCES

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177–208.

[D56] Dawes, B. 1956. The Trematoda with special reference to British and other European forms. University Press: Cambridge.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[H04] Haeckel, E. 1899–1904. Kunstformen der Natur. Bibliographisches Institut: Leipzig und Wien.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H01] Huys, R. 2001. Splanchnotrophid systematics: a case of polyphyly and taxonomic myopia. Journal of Crustacean Biology 21 (1): 106–156.

[KC11] Kocot, K. M., J. T. Cannon, C. Todt, M. R. Citarella, A. B. Kohn, A. Meyer, S. R. Santos, C. Schander, L. L. Moroz, B. Lieb & K. M. Halanych. 2011. Phylogenomics reveals deep molluscan relationships. Nature 477: 452–456.

[MG-H11] McEnnulty, F. R., K. L. Gowlett-Holmes, A. Williams, F. Althaus, J. Fromont, G. C. B. Poore, T. D. O’Hara, L. Marsh, P. Kott, S. Slack-Smith, P. Alderslade & M. V. Kitahara. 2011. The deepwater megabenthic invertebrates on the western continental margin of Australia (100–1100 m depths): composition, distribution and novelty. Records of the Western Australian Museum Supplement 80: 1–191.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II pt I. Stanford University Press: Stanford University (California).

[PP78] Poorman, F. L., & L. H. Poorman. 1978. Additional molluscan records from Bahía de Los Angeles, Baja California Norte. Veliger 20 (4): 369–374.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

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