Trochoidea

Stomatia phymotis, copyright Femorale.


Belongs within: Vetigastropoda.
Contains: Anomphalinae, Pycnomphalinae, Solariella, Margaritinae, Trochinae, Umboniinae, Calliostomatinae, Stomatella, Margarella.

The Trochoidea is a diverse group of vetigastropods lacking either a slit or selenizone (Bouchet & Rocroi 2005). In recent forms the operculum, if present, is corneous, in contrast to the calcareous operculum of the Turbinoidea.

<==Trochoidea [Microdomatacea, Microdomatoidea, Trochacea]
    |--Tychobrahea Horný 1992 [Tychobraheidae] BR05
    |--Eucochlis Knight 1933 [Eucochlidae] BR05
    |    `--*E. perminuta Knight 1933 KC60
    |--Velainella Vasseur 1880 BR05 [=Velainiella (l. c.) KC60; Velainellidae, Velainellinae, Vellainelloidea]
    |    `--*V. columnaris Vasseur 1880 KC60
    |--Elasmonematidae BR05
    |    |--Elasmonema Fischer 1885 BR05 [=Callonema Hall 1879 non Conrad 1875 KC60]
    |    |    `--*E. bellatulum (Hall 1861) [=Loxonema bellatulum, *Callonema bellatulum] KC60
    |    |--Anematina Knight 1933 B79
    |    |    `--*A. proutana (Hall 1858) B79 [=Holopea proutana KC60]
    |    |--Streptotrochus Perner 1907 KC60
    |    |    `--*S. rugulosus Perner 1907 KC60
    |    `--Discordichilus Cossmann 1918 KC60
    |         `--*D. mollis (Lindström 1884) [=Trochus mollis] KC60
    |--Anomphalidae [Anomphalacea] P84
    |    |  i. s.: Cycloscena Fletcher 1958 W63
    |    |           |--*C. anomphala Fletcher 1958 W63
    |    |           |--C. costata Waterhouse 1963 W63
    |    |           |--C. glabra Waterhouse 1963 W63
    |    |           `--‘Pleurotomaria’ rotunda Etheridge 1872 [=Platyschisma rotunda] W63
    |    |         Eiselia Dietz 1911 P84
    |    |           `--*E. dyadica Dietz 1911 KC60
    |    |--Anomphalinae P84
    |    `--Pycnomphalinae P84
    |--Solariellidae [Solariellinae] BR05
    |    |--Solariella BR05
    |    |--Cidarina Dall 1909 KC60
    |    |    `--*C. cidaris (Carpenter 1864) KC60 [=Margarita cidaris KC60, Margarites cidaris O27]
    |    `--Minolia Adams 1860 H92 [=Minosia Dunker 1882 KC60; incl. Minolops Iredale 1929 KC60; Minoliinae]
    |         |--*M. punctata Adams 1860 H92
    |         |--M. casta (Nevill & Nevill 1874) H92
    |         |--M. gradata Sowerby 1895 H92
    |         |--M. holdsworthana Nevill & Nevill 1871 H92
    |         |--M. nedyma Melvill 1897 BD86
    |         |--M. singaporensis (Pilsbry 1889) [=Monilea (Minolia) singaporensis] BC01
    |         |--M. strigata Tenison-Woods 1878 F71
    |         `--M. vitiliginea (Menke 1843) H92
    |--Holopeidae [Holopeinae] PE02
    |    |--Raphispira Perner 1903 KC60
    |    |    `--*R. plena Perner 1903 KC60
    |    |--Cyclora Hall 1845 [Cycloridae] BR05
    |    |    `--*C. minuta Hall 1845 KC60
    |    |--Straparollina Billings 1865 [Straparollinidae, Straparollinoidea] KC60
    |    |    `--*S. pelagica Billings 1865 KC60
    |    |--Globonema Wenz 1938 KC60
    |    |    `--*G. kokeni (Wenz 1938) (see below for synonymy) KC60
    |    |--Protospirialis Clarke 1904 KC60
    |    |    `--*P. minutissima (Clarke 1885) [=Platyostoma minutissima] KC60
    |    `--Holopea Hall 1847 BR05 (see below for synonymy)
    |         |--*H. symmetrica Hall 1847 PE02 [=*Litiopsis symmetrica KC60]
    |         |--H. baltica YN83
    |         |--H. rotunda YN83
    |         |--H. ruoqiangensis Yu & Ning 1983 YN83
    |         |--H. teres Pan & Erwin 2002 PE02
    |         `--H. wellingtonensis Etheridge 1898 F71
    |--Proconulidae [Proconulinae] BR05
    |    |--Dimorphotectus Cossmann 1918 KC60
    |    |    `--D. fasciatus (Hoernes 1856) (see below for synonymy) KC60
    |    |--Callotrochus Kutassy in Wenz 1938 [=Mesotrochus Kutassy 1927 non Wasmann 1890] KC60
    |    |    `--*C. triadicus (Kutassy 1927) [=*Mesotrochus triadicus] KC60
    |    |--Eocalliostoma Haas 1953 KC60
    |    |    `--*E. interruptum (Cox 1947) [=Calliostoma interruptum] KC60
    |    |--Epulotrochus Cossmann 1918 KC60
    |    |    `--*E. epulus (d’Orbigny 1850) [=Trochus epulus] KC60
    |    |--Lithotrochus Conrad 1855 KC60
    |    |    `--L. humboldtii (von Buch 1839) (see below for synonymy) KC60
    |    |--Trypanotrochus Cossmann 1918 KC60
    |    |    `--*T. normanianus (d’Orbigny 1850) [=Trochus normanianus] KC60
    |    |--Proconulus Cossmann 1918 BR05
    |    |    |--*P. guillieri (Cossmann 1885) [=Trochus guillieri] KC60
    |    |    `--P. raulineus KC60
    |    |--Diplochilus Wöhrmann 1894 [=Raiblia Cossmann 1916, Woehrmannia Cossmann 1916] KC60
    |    |    |--*D. gracilis Wöhrmann 1894 [=*Raiblia gracilis, *Woehrmannia gracilis] KC60
    |    |    `--D. bistriatus KC60
    |    |--Tylotrochus Koken 1896 KC60
    |    |    |--*T. konincki (Hörnes 1856) [=Trochus konincki] TE01
    |    |    `--T. elongatus Bandel 1993 [=Ziziphinus semipunctatus Zardini 1978 (n. n.)] TE01
    |    |--Anticonulus Cossmann 1918 KC60
    |    |    |--*A. mariae (d’Orbigny 1853) [=Trochus mariae] KC60
    |    |    `--A. nisus KC60
    |    |--Muricotrochus Cossmann 1918 KC60
    |    |    |--*M. hudlestoni Cossmann 1918 KC60
    |    |    `--M. subluciensis KC60
    |    `--Discotectus Favre 1913 KC60
    |         |--*D. massalongoi [=Trochus massalongoi] KC60
    |         `--D. crassiplicatus KC60
    |--Microdomatidae BR05
    |    |  i. s.: Episfaxis Perner 1903 FB04 [=Cosmina Perner 1903 non Robineau-Desvoidy 1830 KC60]
    |    |           `--*E. complacens (Perner 1903) [=*Cosmina complacens] KC60
    |    |         Glyptospira Chronic 1952 FB04
    |    |           |--*G. cristulata Chronic 1952 PE02
    |    |           `--G. pinna Pan & Erwin 2002 PE02
    |    |         Daidia Wilson 1951 FB04
    |    |           `--*D. cerithioides (Salter 1859) [=Eunema cerithioides] KC60
    |    |         Pagodea Perner 1903 FB04
    |    |           `--*P. concomitans Perner 1903 KC60
    |    |         Dongiovannia Horný 1992 FB04
    |    |         Petrochus Horný 1992 FB04
    |    |         Threavia Lamont 1946 FB04
    |    |           `--*T. gulosa Lamont 1946 KC60
    |    |         Eopagodea Frýda & Rohr 2001 FB04
    |    |--Microdoma Meek & Worthen 1866 BR05 (see below for synonymy)
    |    |    |--*M. conicum Meek & Worthen 1867 FB04
    |    |    |--M. bicrenulata (De Koninck 1883) B79
    |    |    |--M. biserratum (Phillips 1836) B79
    |    |    |--M. nodosum Batten 1979 B79
    |    |    |--M. serrilimba (Phillips 1836) B79
    |    |    `--M. variegata Batten 1979 B79
    |    `--Decorospirinae BR05
    |         |--Dutrochus Blodgett 1993 FB04
    |         |--Roemeriella Blodgett & Frýda 1999 FB04
    |         `--Decorospira Blodgett & Johnson 1992 BR05
    |              |--*D. tasselli Blodgett & Johnson 1992 FB04
    |              |--D. gilberti Blodgett & Johnson 1992 FB04
    |              |--D. lepaini Frýda & Blodgett 2004 FB04
    |              |--D. minutula Frýda & Blodgett 2004 FB04
    |              |--D. pragensis (Horný 1992) FB04
    |              `--D. rigbyi Blodgett & Johnson 1992 FB04
    `--Trochidae [Calliostomatidae, Trochinia] BR05
         |  i. s.: Cirsostylus Cossmann 1918 TE01, KC60
         |           `--*C. glandulus (Laube 1868) [=Trochus glandulus] TE01
         |         Kittlitrochus Cossmann 1909 (see below for synonymy) KC60
         |           `--*K. marginenodosus (Böhm 1895) (see below for synonymy) KC60
         |         Trochodon Seeley 1861 KC60
         |           `--*T. cancellatus (Seeley 1861) [=Trochus (*Trochodon) cancellatus] KC60
         |--Margaritinae H92
         |--Trochinae H92
         |--Umboniinae H92
         |--Calliostomatinae S11
         |--Thysanodonta Marshall 1988 [Thysanodontidae, Thysanodontinae] BR05
         |--Halistylus Dall 1890 [Halistylinae] BR05
         |    |--*H. columna (Dall 1890) [=Cantharidus (*Halistylus) columna] KC60
         |    `--H. subpupoideus Tryon 1887 [=Fenella pupoidea (preoc.)] O27
         `--Stomatellinae [Macrostoma, Stomatellidae, Stomatidae, Stomatiidae, Stomatiinae] BR05
              |--Stomatella BW09
              |--Stylobates F27
              |--Phaneta everetti G88
              |--Margarella P61
              |--Broderipia Gray 1847 KC60
              |    `--*B. rosea (Broderip 1834) [=Scutella rosea] KC60
              |--Mesogena Kutassy ex Cox in Knight, Cox et al. 1960 KC60
              |    `--*M. arcta (Hoernes 1855) [=Inoceramus arctus] KC60
              |--Praestomatia Kutassy ex Cox in Knight, Cox et al. 1060 KC60
              |    `--*P. acutangula (Koken 1897) [=Stomatia acutangula] KC60
              |--Roya Iredale 1912 KC60
              |    `--*R. kermadecensis Iredale 1912 KC60
              |--Synaptocochlea Pilsbry 1890 KC60
              |    |--S. stellata (Souverbie 1863) (see below for synonymy) KC60
              |    `--S. picta (d’Orbigny 1842) [incl. S. picta var. lactea Usticke 1959] BC01
              |--Pseudostomatella Thiele 1924 KC60
              |    |  i. s.: P. erythrocoma BC01
              |    |           |--P. e. erythrocoma BC01
              |    |           `--P. e. cycloradiata Usticke 1959 BC01
              |    |--*P. (Pseudostomatella) papyracea (Gmelin 1791) KC60 (see below for synonymy)
              |    `--P. (Stomatolina Iredale 1937) KC60
              |         `--P. (*S.) rufescens (Gray 1847) [=Stomatella rufescens] KC60
              |--Stomatia Helbling 1779 BR05 [=Stomax Montfort 1810 KC60; incl. Niphonia F27]
              |    |  i. s.: S. angulata Adams 1850 H09
              |    |         S. australis Adams 1850 H09
              |    |         S. decussata Adams 1850 H09
              |    |--*S. (Stomatia) phymotis Helbling 1779 [=*Stomax phymotis] KC60
              |    |--S. (Microtis Adams & Angas 1850 non Br. 1810 [ICBN]) KC60
              |    |    `--S. (*M.) tuberculata (Adams 1850) [=*Microtis tuberculata] KC60
              |    `--S. (Miraconcha Bergh in Schepman 1908) KC60
              |         `--S. (*M.) obscura (Bergh in Schepman 1908) [=*Miraconcha obscura] KC60
              `--Herpetopoma Pilsbry 1889 P61
                   |--*H. scabriuscula (Adams & Angas 1867) [=Euchelus scabriusculus] KC60
                   |--H. aspersa M54 [incl. Euchelus baccatus F27]
                   |--H. bella (Hutton 1873) [=Euchelus (Herpetopoma) bellus; incl. E. hamiltoni (Kirk. 1882)] F27
                   |--H. benthicola Powell 1937 P61
                   |--H. larochei Powell 1926 [=Euchelus (Herpetopoma) larochei] F27
                   |    |--H. l. larochei P61
                   |    `--H. l. alacerrima Dell 1956 P61
                   |--H. mariae Finlay 1930 P61
                   `--H. scaber M54

Dimorphotectus fasciatus (Hoernes 1856) [=Scoliostoma fasciatum; incl. Tectus hoernesi Koken 1896, *Dimorphotectus hoernesi] KC60

*Globonema kokeni (Wenz 1938) [=Trochonema (*Globonema) kokeni, Nematotrochus bicarinatus Koken 1925 non (Wahlenberg 1821)] KC60

Holopea Hall 1847 BR05 [=Litiopsis Salter 1866 KC60; incl. Anastrophina Knight 1937 KC60, Cirropsis Perner 1903 KC60, Haplospira Koken 1897 KC60, Staurospira Perner 1907 non Haeckel 1887 KC60, Tortilla Perner 1903 KC60]

Kittlitrochus Cossmann 1909 [=Paratrochus Kittl 1899 non Pilsbry 1893, Paratrochoides Tomlin 1929] KC60

*Kittlitrochus marginenodosus (Böhm 1895) [=Tectus marginenodosus, *Paratrochoides marginenodosus, *Paratrochus marginenodosus] KC60

Lithotrochus humboldtii (von Buch 1839) [=Pleurotomaria humboldtii; incl. Turritella andii d’Orbigny 1842, *Lithotrochus andii] KC60

Microdoma Meek & Worthen 1866 BR05 [=Microdomus Cossmann 1915 KC60; incl. Tuberculopleura Jakowlew 1899 KC60; Microdomatinae, Microdominae]

*Pseudostomatella (Pseudostomatella) papyracea (Gmelin 1791) KC60 [=Turbo papyraceus KC60, Stomatella papyracea KC60, Stomatia (*Pseudostomatella) papyracea F27]

Synaptocochlea stellata (Souverbie 1863) [=Stomatella stellata; incl. St. montrouzieri Pilsbry 1890, *Synaptocochlea montrouzieri] KC60

*Type species of generic name indicated

REFERENCES

[BD86] Barash, A., & Z. Danin. 1986. Further additions to the knowledge of Indo-Pacific Mollusca in the Mediterranean Sea (Lessepsian migrants). Spixiana 9 (2): 117–141.

[B79] Batten, R. L. 1979. Gastropods from Perak, Malaysia. Part 2. The trochids, patellids and neritids. American Museum Novitates 2685: 1–26.

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[BW09] Bryce, C., & C. Whisson. 2009. The macromolluscs of Mermaid (Rowley Shoals), Scott and Seringapatam Reefs, Western Australia. Records of the Western Australian Museum Supplement 77: 177–208.

[F27] Finlay, H. J. 1927. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F71] Fletcher, H. O. 1971. Catalogue of type specimens of fossils in the Australian Museum, Sydney. Australian Museum Memoir 13: 1–167.

[FB04] Frýda, J., & R. B. Blodgett. 2004. New Emsian (Late Early Devonian) gastropods from Limestone Mountain, Medfra B-4 quadrangle, west-central Alaska (Farewell Terrane), and their paleobiogeographic affinities and evolutionary significance. Journal of Paleontology 78 (1): 111–132.

[G88] Gray, J. 1988. Evolution of the freshwater ecosystem: the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology 62: 1–214.

[H09] Hedley, C. 1909. The Marine Fauna of Queensland: Address by the President of Section D. Australasian Association for the Advancement of Science: Brisbane.

[H92] Herbert, D. G. 1992. Revision of the Umboniinae in southern Africa and Mozambique (Mollusca: Prosobranchia: Trochidae). Annals of the Natal Museum 33 (2): 379–459.

[KC60] Knight, J. B., L. R. Cox, A. M. Keen, R. L. Batten, E. L. Yochelson & R. Robertson. 1960. Gastropoda: systematic descriptions. In: Moore, R. C. (ed.) Treatise on Invertebrate Paleontology pt I. Mollusca 1: Mollusca—General Features, Scaphopoda, Amphineura, Monoplacophora, Gastropoda—General Features, Archaeogastropoda and some (mainly Paleozoic) Caenogastropoda and Opisthobranchia pp. I169–I331. Geological Society of America, and University of Kansas Press.

[M54] Macpherson, J. H. 1954. Molluscs (sea shells and snails). In: Willis, J. M. (ed.) The Archipelago of the Recherche pp. 55–63. Australian Geographical Society: Melbourne.

[O27] Oldroyd, I. S. 1927. The Marine Shells of the West Coast of North America vol. II pt. III. Stanford University Press: Stanford University (California).

[PE02] Pan, H.-Z., & D. H. Erwin. 2002. Gastropods from the Permian of Guangxi and Yunnan Provinces, south China. Journal of Paleontology 76 (Memoir 56): 1–49.

[P84] Peel, J. S. 1984. Autecology and systematics of a new Silurian anomphalid gastropod from western North Greenland. Rapp. Grønlands geol. Unders. 121: 77–87.

[P61] Powell, A. W. B. 1961. Shells of New Zealand: An illustrated handbook 4th ed. Whitcombe and Tombs Limited: Christchurch.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

[TE01] Tong, J., & D. H. Erwin. 2001. Triassic gastropods of the southern Qinling Mountains, China. Smithsonian Contributions to Paleobiology 92: 1–47.

[W63] Waterhouse, J. B. 1963. Permian gastropods of New Zealand. Part 4—Platyceratacea, Anomphalacea, Neritacea, and correlations. New Zealand Journal of Geology and Geophysics 6 (5): 817–842.

[YN83] Yu W. & Ning H. 1983. Middle Ordovician gastropods from Ruoqiang, Xinjiang. Acta Palaeontologica Sinica 22 (2): 199–203.

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