Cancellariidae

Plesiotriton vivus, copyright MNHN, Paris.


Belongs within: Siphonogastropoda.
Contains: Unitas, Zeadmete, Oamaruia, Inglisella, Bonellitia, Merica, Sveltella, Tritonoharpa, Turehua, Sveltia, Cancellaria, Trigonostoma.

The Cancellariidae, nutmeg snails, are a group of marine gastropods adapted for suctorial feeding, found in subtidal to bathyal sand and mud bottoms (Petit & Harasewych 1990). Among the subfamilies of the Cancellariidae, members of the Plesiotritoninae have more elongate shells than the others. The Cancellariinae have thicker shells with columellar pleats, whereas Admetinae have thin shells that usually lack columellar pleats.

Synapomorphies (from Petit & Harasewych 1990): Jaw anteriorly tubular, posteriorly saddle-shaped, spanning dorsal surface of buccal mass; radula uniserial, composed of extremely elongated (length/width ratio >15), ribbon-like, tricuspid teeth with secondary dentition on one or more cusps (radula sometimes absent); anterior esophagus extremely short, valve of Leiblein and buccal ganglia situated at posterior margin of buccal mass; gland of Leiblein absent.

Cancellariidae [Cancellariacea, Cancellariadae, Cancellarioidea, Cancellinae]
    |--Cancellariinae BR05
    |    |  i. s.: Sveltia O27a
    |    |         Crawfordina Dall 1919 O27a, PH90 [=Crawfordia Dall 1918 non Pierce 1908 PH90]
    |    |           `--*C. crawfordiana (Dall 1891) (see below for synonymy) PH90
    |    |--Cancellaria S11
    |    `--+--Nevia Jousseaume 1887 PH90
    |       |    `--*N. spirata (Lamarck 1822) PH90, S11 [=Cancellaria spirata S11; incl. C. excavata Sowerby 1849 PH90]
    |       `--+--Trigonostoma S11
    |           `--Scalptia Jousseaume 1887 S11, PH90
    |                |--*S. obliquata (Lamarck 1822) (see below for synonymy) PH90
    |                |--S. contabulata (Sowerby 1833) S11
    |                |--S. hystrix (Reeve 1856) (see below for synonymy) PH90
    |                |--S. mercadoi Old 1968 PH90
    |                |--S. nassa (Gmelin 1791) [=Voluta nassa; incl. Cancellaria lamellosa Hinds 1843] PH90
    |                `--S. scala (Gmelin 1791) (see below for synonymy) PH90
    |--Plesiotriton Fischer 1884 [Plesiotritoninae] BR05
    |    |--*P. volutella (Lamarck 1803) [=Cancellaria volutella, C. volvutella (l. c.)] PH90
    |    |--P. cedri (Gardner 1935) [=Tritonium cedri, T. ceden (l. c.)] PH90
    |    |--P. chalmasi (Quass 1902) [=Triton chalmasi, T. chalmsi (l. c.)] PH90
    |    |--P. cossmanni Oppenheim 1906 PH90
    |    |--P. cretaceus Sohl 1960 PH90
    |    |--P. deshayesianus Beu & Maxwell 1987 (see below for synonymy) PH90
    |    |--P. farafrense (Quaas 1902) [=Triton farafrense] PH90
    |    |--P. hillegondae (Martin 1914) [=Eutritonium hillegondae] PH90
    |    |--P. imbricatum (Dareste de la Chavanne 1910) [=Tritonium imbricatum] PH90
    |    |--P. lanceolata (Menke 1828) [=Colubraria leona Mansfield 1937] PH90
    |    |--P. mirabilis Beu & Maxwell 1987 PH90
    |    |--P. multicostatum (Favre 1869) [=Triton multicostatum] PH90
    |    |--P. paytensis Olsson 1930 PH90
    |    |--P. vivus Habe & Okutani 1981 PH90
    |    `--P. vokesae (Allen 1970) [=Colubraria vokesae] PH90
    `--Admete Möller 1842 [Admetacea, Admetidae, Admetinae] BR05
         |--A. viridula (Fabricius 1780) (see below for synonymy) PH90
         |    |--A. v. viridula PH90
         |    |--A. ‘couthouyi’ gracilior Carpenter 1866 O27a
         |    `--A. v. sheppardi Bell 1919 PH90
         |--A. aethiopica Thiele 1925 PH90
         |--A. antarctica Strebel 1908 PH90
         |--A. beatrix Olsson 1964 PH90
         |--A. azorica Bouchet & Warén 1985 PH90
         |--A. biplicata (Ravn 1902) [=Tritonium biplicatum] PH90
         |--A. bruuni Knudsen 1964 PH90
         |--A. californica Dall 1908 O27a
         |--A. cancellata Kobelt 1887 PH90
         |--A. chainei Peyrot 1928 PH90
         |--A. choshiensis Shikama 1962 PH90
         |--A. clatskaniensis Anderson & Martin 1914 PH90
         |--A. consobrina Powell 1951 PH90
         |--A. cornidei Altimira 1978 PH90
         |--A. couturieauxi Glibert 1938 PH90
         |--A. decapensis Barnard 1960 PH90
         |--A. delicatula Smith 1907 PH90
         |--A. emmae Albrecht & Valk 1943 PH90
         |--A. enderbyensis Powell 1958 PH90
         |--A. exilis [incl. A. exilis var. angulopusilla Sacco 1894, A. exilis var. angulosior Sacco 1804] PH90
         |--A. frigida Rochebrune & Mabille 1885 PH90
         |--A. funigera Staadt in Cossmann 1913 PH90
         |--A. girauxi Cossmann 1913 PH90
         |--A. harpovoluta Powell 1957 PH90
         |--A. hukuiensis Nomura & Niino 1940 PH90
         |--A. laevior Leche 1878 PH90 (see below for synonymy)
         |--A. latecostata Traub 1938 PH90
         |--A. lirisculpta Cossmann 1902 PH90
         |--A. luffa Olsson 1929 PH90
         |--A. magellanica Strebel 1905 PH90
         |--A. maorium Marshall & Murdoch 1921 PH90
         |--A. microsoma Dall 1908 O27a
         |--A. minuta (see below for synonymy) PH90
         |--A. nassiformis (see below for synonymy) PH90
         |--A. nodosa Verrill & Smith in Verrill 1885 PH90
         |--A. oligopercostata Sacco 1894 PH90
         |--A. olssoni Rivera 1957 PH90
         |--A. ornata Il’ina 1955 PH90
         |--A. ovata Smith 1875 PH90
         |--A. paytensis Olsson 1930 PH90
         |--A. philippi von Ihering 1907 [=Cancellaria australis Philippi 1855 non Sowerby 1832] PH90
         |--A. reedii Bell 1870 PH90
         |--A. regina Dall 1911 PH90
         |--A. rhyssa Dall 1919 PH90
         |--A. sadko Gorbunov 1946 PH90
         |--A. seftoni Berry 1956 PH90
         |--A. sphaeroidea Luković 1924 PH90
         |--A. subfusiformis Meek 1873 PH90
         |--A. submitroides Cossmann & Pissarro 1905 PH90
         |--A. tabulata Sowerby 1875 PH90
         |--A. triplicata Namias 1898 PH90
         |--A. umbilicata Hickman 1980 PH90
         |--A. unalashkensis (Dall 1873) PH90 (see below for synonymy)
         |--A. veneranda Beets 1946 PH90
         |--A. watanabei Shikama 1962 PH90
         |--A. woodii Sacco 1894 PH90
         |--A. woodworthi Dall 1905 PH90
         |--A. wouweri Harmer 1918 PH90
         `--A. yokoyamai Oyama 1954 [=Lora cancellata Otuka 1937 non Admete cancellata Kobelt 1887] PH90

Cancellariidae incertae sedis:
  Aphera Adams & Adams 1854 PH90
    |--*A. tessellata (Sowerby 1832) [=Cancellaria tessellata] PH90
    |--A. bronni (see below for synonymy) PH90
    |--A. exdecussata Sacco 1894 PH90
    |--A. lindae Petuch 1987 PH90
    |--A. ovatocrassa Sacco 1894 PH90
    |--A. scopalveus Finlay 1926 PH90
    `--A. waltonensis (Aldrich 1910) [=Mitra waltonensis] PH90
  Unitas PH90
  Maorivetia Finlay 1924 PH90
    `--*M. brevirostris (Hutton 1877) [=Turbinella brevirostris] PH90
  Microsveltia Iredale 1925 PH90
    `--*M. recessa Iredale 1925 PH90
  Zeadmete PH90
  Pepta Iredale 1925 PH90
    |--*P. stricta (Hedley 1907) [=Admete stricta] PH90
    `--P. turriculata F27a
  Oamaruia PH90
  Inglisella PH90
  Bivetia Jousseaume 1887 PH90
    |--‘Cancellaria’ indentata Sowerby 1832 (see below for synonymy) PH90
    |--B. martini Cossmann 1899 (see below for synonymy) PH90
    `--B. neuvillei Peyrot 1928 PH90
  Admetopsis Meek 1873 PH90
    |--*A. gregaria (Meek 1873) [=Admete gregaria] PH90
    |--A. elevata Johnson 1903 PH90
    |--A. humerosa Stanton 1893 PH90
    `--A. rhomboides (Meek 1873) [=Admete rhomboides] PH90
  Admetula Cossmann 1889 PH90
    |--*A. evulsa (Solander 1766) (see below for synonymy) PH90
    |--A. bayeri Petit 1976 PH90
    |--A. deroyae (Petit 1970) [=Agatrix deroyae] BC01
    |--A. garrardi Petit 1974 [=Cancellaria nassoides Schepman 1911 non von Koenen 1889] PH90
    |--A. inflata Dockery in MacNeil & Dockery 1984 PH90
    |--A. paucivaricata (Gabb 1864) [=Tritonium paucivaricatum] PH90
    |--A. regularia Dockery in MacNeil & Dockery 1984 PH90
    `--A. vossi Petit 1976 PH90
  Africostoma Eames 1957 PH90
    `--*A. decorata (Newton 1922) [=Trigonostoma decorata] PH90
  Africosveltia Eames 1957 PH90
    `--*A. multiplicis (Newton 1922) [=Cancellaria multiplicis] PH90
  Africotriton Beu & Maxwell 1987 PH90
    |--*A. crebriliratus (Sowerby 1903) [=Epidromus crebriliratus] PH90
    |--A. carinapex Beu & Maxwell 1987 PH90
    |--A. fictilis Hinds 1844 PH90
    |--A. kilburni Beu & Maxwell 1987 PH90
    `--A. multinodulatus Beu & Maxwell 1987 PH90
  Agatrix Petit 1967 PH90
    |--*A. agassizii (Dall 1889) [=Trigonostoma agassizii] PH90
    `--A. nodosivaricosa Petuch 1979 PH90
  Anapepta Finlay 1930 PH90
    |--*A. anomala (Marshall & Murdoch 1920) [=Admete anomala] PH90
    |--A. explicata Laws 1935 PH90
    |--A. finlayi Marwick 1931 PH90
    |--A. nucleosa Marwick 1931 PH90
    |--A. serrata Laws 1935 PH90
    `--A. tuberculifera Laws 1935 PH90
  Aneurystoma Cossmann 1899 PH90
    `--*A. dufourii (Grateloup 1832) (see below for synonymy) PH90
  Antepepta Finlay & Marwick 1937 PH90
    `--*A. nasuta Finlay & Marwick 1937 PH90
  Arizelostoma Iredale 1936 PH90
    `--*A. laseroni Iredale 1936 PH90
  Axelella Petit 1988 [=Olssonella Petit 1970 non Glibert & Van de Poel 1967] PH90
    |--*A. smithii (Dall 1888) [=Cancellaria smithii, *Olssonella smithii] PH90
    |--‘Olssonella’ elongata Dockery in MacNeil & Dockery 1984 PH90
    |--‘Olssonella’ panamica Petit 1976 PH90
    |--‘Olssonella’ vokesae Petit 1970 PH90
    `--‘Olssonella’ williamsi Petit 1976 PH90
  Babylonella Conrad 1865 PH90
    |--*B. elevata (Lea 1833) [=Cancellaria elevata] PH90
    |--B. nassiformis PH90
    |    |--B. n. nassiformis PH90
    |    `--B. n. profunda Tabanelli 1985 PH90
    |--B. pusilla (Philippi 1843) (see below for synonymy) PH90
    `--B. stemerdinki Janse & Janssen 1983 PH90
  ‘Barkeria’ Addicott 1970 non Szidat 1936 PH90
    `--*B. sanjosei (Anderson & Martin 1914) [=Cancellaria sanjosei, C. sanjoseensis] PH90
  Bivetiella Wenz 1943 [=Bivetia Jousseaume 1887 non Jousseaume 1887] PH90
    `--*B. similis (Sowerby 1833) (see below for synonymy) PH90
  Bivetopsia Jousseaume 1887 [=Bivetopsis Cossmann 1899] PH90
    `--*B. chrysostoma (Sowerby 1832) (see below for synonymy) PH90
  Bonellitia PH90
  Brocchinia Jousseaume 1887 PH90
    |--B. parvula [incl. B. parvula var. pluricarinata Sacco 1894] PH90
    |    |--B. p. parvula PH90
    |    `--B. p. tauroparva Sacco 1894 (see below for synonymy) PH90
    |--B. clenchi Petit 1986 [incl. Cancellaria pusilla Adams 1869 non Sowerby 1832] PH90
    |--B. crassinodosa Sacco 1894 PH90
    |--B. marshalli (Vignal in Cossmann 1921) PH90 (see below for synonymy)
    |--B. pyramidatula Sacco 1894 PH90
    `--B. rissoiaeformis Cossmann 1899 PH90
  Calcarata Jousseaume 1887 PH90
    `--*C. calcarata (Brocchi 1814) [=Voluta calcarata] PH90
  Cancellaphera Iredale 1930 PH90
    |--*C. amasia Iredale 1930 PH90
    `--C. confirmans Ludbrook 1958 PH90
  Cancellariella Martin 1904 PH90
    `--*C. neritoidea (Martin 1904) [=Cancellaria (*Cancellariella) neritoidea] PH90
  Cancelrana Palmer 1937 PH90
    |--*C. finexa (Harris 1895) [=Pleurotoma (Taranis) finexa] PH90
    `--C. lischkei PH90
         |--C. l. lischkei PH90
         `--C. l. pauciplicata Shuto 1962 PH90
  Charcolleria Olsson 1942 PH90
    |--*C. perdiciana (Olsson 1942) [=Cancellaria (Charcolleria) perdiciana] PH90
    `--C. distinguenda Petit 1970 PH90
  Contortia Sacco 1894 PH90
    `--*C. contorta (Basterot 1825) (see below for synonymy) PH90
         |--C. c. contorta PH90
         `--‘Cancellaria’ c. gelriana Janssen 1972 PH90
  Coptostoma Cossmann 1899 PH90
    |--*C. quadratum (Sowerby 1822) (see below for synonymy) PH90
    |--C. breve Wrigley 1935 PH90
    |--C. punjabense Eames 1952 PH90
    |--C. rahkiense Eames 1952 PH90
    |--C. rameum Palmer 1944 PH90
    `--C. ulmulum [incl. C. ulmulum var. secutorum Palmer 1947] PH90
  Coptostomella Finlay & Marwick 1937 PH90
    `--*C. pupa Finlay & Marwick 1937 PH90
  Daguinia Magne 1966 PH90
    `--*D. vigneauxi Magne 1966 PH90
  Dellina Beu 1970 PH90
    `--D. aoteana (Dell 1956) [=Antizafra aoteana; incl. Waipaoa munida Ponder 1968, *Dellina munida] PH90
  Egerea Gábor 1936 PH90
    `--*E. collectiva Gábor 1936 [incl. E. collectiva var. nassaeformis Noszky 1936] PH90
  Extractrix Korobkov 1955 PH90
    `--*E. extractrix (Boettger 1906) PH90 [=Discohelix extractrix, Pseudomalaxis extractrix] PH90
  Fusiaphera Habe 1961 PH90
    |--*F. macrospira (Adams & Reeve 1850) [=Cancellaria macrospira] PH90
    |--F. azumai Habe 1961 PH90
    |--F. dampierensis Garrard 1975 PH90
    |--F. eva Petit 1980 PH90
    |--F. macrospiratoides Habe 1961 PH90
    `--F. tosaensis Habe 1961 PH90
  Gerdiella Olsson & Bayer 1972 PH90
    |--*G. gerda Olsson & Bayer 1972 PH90
    |--G. cingulata Olsson & Bayer 1972 PH90
    `--G. santa Olsson & Bayer 1972 PH90
  Gergovia Cossmann 1899 PH90
    |--‘Cancellaria’ laticostata Tenison-Woods 1880 (see below for synonymy) PH90
    `--G. haswelli Garrard 1975 PH90
  Gulia Jousseaume 1887 PH90
    `--*G. acutangula (Faujas de Saint Fond 1817) (see below for synonymy) PH90
  Habesolatia Kuroda 1965 PH90
    `--*H. nodulifera (Sowerby 1825) [=Cancellaria nodulifera, C. noduliformis (l. c.)] PH90
  Hertleinia Marks 1949 PH90
    `--*H. mitriformis (Sowerby 1832) (see below for synonymy) PH90
  ‘Inermia’ Korobkov 1955 (preoc.) PH90
    `--*I. inermis (Pusch 1837) (see below for synonymy) PH90
  Iphinopsis Dall 1924 [incl. Iphinoella Habe 1958] PH90
    |--*I. kelseyi (Dall 1908) [=Trichotropis kelseyi, Iphinoe kelseyi] PH90
    |--I. alba Bouchet & Warén 1985 PH90
    |--I. bathyalis (Okutani 1964) [=Palaeadmete (l. c. for Paladmete) bathyalis] PH90
    |--*Iphinoella’ choshiensis Habe 1958 PH90
    |--I. fuscoapicata Bouchet & Warén 1985 PH90
    |--I. inflata (Friele 1879) [=Trichotropis inflata] PH90
    |--I. kroyeri (Philippi 1849) PH90 [=Trichotropis (Iphinoe) kroyeri PH90, O27b]
    |--I. nuda (Dall 1927) [=Trichotropis nuda] PH90
    `--I. traverseensis (Clarke 1961) [=Tromina traverseensis, T. traversensis (l. c.)] PH90
  Junghuhnia Martin 1931 PH90
    `--*J. ficus (Martin 1931) [=Trigonostoma (*Junghuhnia) ficus] PH90
  Kapuatriton Beu & Maxwell 1987 PH90
    `--*K. kaitarus Beu & Maxwell 1987 PH90
  Loxotaphrus Harris 1897 PH90
    |--*L. variciferus (Tate 1888) [=Phos variciferus] PH90
    |--L. aturensis (Peyrot 1926) [=Tritonidea aturensis] PH90
    |--L. deshayesii (Duval 1841) [=Cassidaria deshayesii] PH90
    |--L. minbuensis (Noetling 1895) [=Cassidaria minbuensis] PH90
    |--L. subtaurinensis (Vergneau 1965) [=Tritonidea subtaurinensis] PH90
    `--L. taurinensis (Bellardi 1873) [=Pollia taurinensis] PH90
  Marianarona Petuch 1988 PH90
    |--*M. alternata (Conrad 1834) [=Cancellaria alternata] PH90
    |--M. asheri Petuch 1988 PH90
    `--M. marylandica Petuch 1988 PH90
  Mariasveltia Petuch 1988 PH90
    `--*M. lunata (Conrad 1830) (see below for synonymy) PH90
  Marksella Olsson 1964 PH90
    `--*M. jumala (Olsson 1964) [=Admete (*Marksella) jumala] PH90
  Massyla Adams & Adams 1854 PH90
    `--*M. corrugata (Hinds 1843) [=Cancellaria corrugata] PH90
  Merica PH90
  Mericella Thiele 1929 PH90
    |--*M. jucunda (Thiele 1925) [=Cancellaria jucunda] PH90
    `--M. bozzettii Petit & Harasewych 1993 BC01
  Microcancilla Dall 1924 PH90
    `--*M. microscopica (Dall 1889) [=Cancellaria microscopica, Admete microscopica] PH90
  Misteia Janssen 1984 PH90
    `--*M. planispira (Nyst 1845) (see below for synonymy) PH90
  Momoebora Kuroda & Habe 1971 PH90
    `--*M. sinensis (Reeve 1856) [=Cancellaria sinensis] PH90
  Narona Adams & Adams 1854 [=Panarona Petit 1975] PH90
    |--*N. clavatula (Sowerby 1832) (see below for synonymy) PH90
    |--N. canaliculata Janssen 1972 PH90
    |--N. dingdensis Anderson 1964 PH90
    |--N. eximia Stoliczka 1867
    |--N. fusiformis [incl. N. fusiformis var. crassicostata Nordsieck 1972] PH90
    |--N. mitraeformis POH88
    |--N. ravni Glibert 1960 [=Cancellaria angulifera von Koenen 1885 non Deshayes 1864] PH90
    `--N. tricarinata Traub 1979 PH90
  Neadmete Habe 1961 PH90
    |--*N. okutanii Petit 1974 PH90
    |--N. nakayamai Habe 1961 PH90
    |--N. nausorensis Ladd 1982 [incl. Hindsia rewaensis Ladd 1982] PH90
    |--N. profundicola Okutani 1964 PH90
    |--N. sagamiensis Kuroda & Habe 1971 PH90
    |--N. schwartzi Mount 1970 PH90
    `--N. sutherlandi Kanakoff & McLean 1966 PH90
  Nipponaphera Habe 1961 PH90
    |--*N. habei Petit 1972 PH90
    `--N. iwaotakii Habe 1961 PH90
  Nothoadmete Oliver 1982 PH90
    `--*N. tumida Oliver 1982 PH90
  Ovilia Jousseaume 1887 PH90
    |--*O. doliolaris (Basterot 1825) (see below for synonymy) PH90
    |--O. fenollerae Landau 1984 PH90
    `--O. onubensis Landau 1984 PH90
  Palaeocancellaria Kollmann 1976 PH90
    `--*P. hoelleitenensis Kollmann 1976 PH90
  Pallidonia Laseron 1955 PH90
    `--*P. simplex Laseron 1955 PH90
  Perplicaria Dall 1890 PH90
    |--*P. perplexa Dall 1890 PH90
    |--P. clarki Smith 1947 PH90
    `--P. prior Maury 1910 PH90
  Pyruclia Olsson 1932 [=Peruclia (l. c.)] PH90
    `--*P. solida (Sowerby 1832) [=Cancellaria solida] PH90
  Plesiocerithium Cossmann 1889 PH90
    `--*P. maglorii (Melleville 1843) (see below for synonymy) PH90
  Pristimerica Finlay & Marwick 1937 PH90
    `--*P. dolioides Finlay & Marwick 1937 PH90
  Rhomboidestoma Martin 1931 PH90
    `--*R. oscitans Martin 1931 [=Trigonostoma orcitans (l. c.)] PH90
  Semitriton Cossmann 1903 PH90
    |--*S. dennanti (Tate 1898) [=Plesiotriton dennanti] PH90
    |--S. inopinatus Cossmann & Pissarro 1905 PH90
    |--S. transennum [=Cymatium transennum] F27a
    `--S. varicosus (Tate 1888) [=Cantharus varicosus] PH90
  Solatia Jousseaume 1887 PH90
    |--S. piscatoria (Gmelin 1791) [=Buccinum piscatorium, *Solatia solat Jousseaume 1887] PH90
    `--S. hirta (Brocchi 1814) [=Voluta hirta] PH90
  Solutosveltia Habe 1961 PH90
    `--*S. abyssicola Habe 1961 PH90
  Sveltella PH90
  Sydaphera Iredale 1929 PH90
    |--‘Cancellaria’ undulata Sowerby 1849 (see below for synonymy) PH90
    |--S. delicosa Laseron 1955 [=S. deliciosa (l. c.)] PH90
    |--S. horii Masuda 1967 PH90
    `--S. obnixa Iredale 1936 PH90
  Tatara Fleming 1950 PH90
    |--*T. pahiensis (Marshall & Murdoch 1921) [=Cymatium pahiense] PH90
    |--T. flemingi Beu & Maxwell 1987 PH90
    |--T. revoluta (Finlay 1924) PH90 [=Cymatium revolutum PH90, Semitriton revolutum F27a]
    `--T. transenna (Suter 1917) [=Ficus transennus] PH90
  Tribia Jousseaume 1887 PH90
    |--*T. angasi (Crosse 1863) [=Cancellaria angasi; incl. C. eudeli Sowerby 1893] PH90
    `--T. epomis Woodring 1928 PH90
  Trigonaphera Iredale 1936 PH90
    |--*T. vinnula (Iredale 1925) [=Trigonostoma vinnulum] PH90
    |--T. amakusana Petit 1974 PH90
    |--T. interlaevis Laseron 1955 PH90
    |--T. stenomphala Habe 1961 PH90
    |--T. teramachii Habe 1961 PH90
    `--T. tosaensis Habe 1961 PH90
  Tritonoharpa PH90
  Turehua PH90
  Varicohilda Eames 1957 PH90
    `--*V. turriculata (Newton 1922) [=Hilda turriculata] PH90
  Vercomaris Garrard 1975 PH90
    `--*V. pergradata (Verco 1904) PH90 [=Cancellaria pergradata PH90, Oamaruia pergradata F27b]
         |--V. p. pergradata PH90
         `--‘Oamaruia’ p. profundior Cotton & Godfrey 1932 PH90
  Waipaoa Marwick 1931 PH90
    |--*W. cristata (Marwick 1926) [=Admete cristata] PH90
    |--W. affectata Marwick 1931 PH90
    |--W. delli Beu 1970 PH90
    |--W. grata Marwick 1931 PH90
    `--W. marwicki Dell 1956 PH90
  ‘Fossarus’ dalli Anderson & Martin 1914 [=F. barkeri Anderson in Hanna 1924] PH90
  ‘Voluta’ benedeni Müller 1851 PH90
  ‘Voluta’ camdeo Forbes 1846 [=V. camdio (l. c.)] PH90
  Pisanella cathalai Doncieux 1908 PH90
  ‘Fasciolaria’ cordensis Stanton 1920 PH90
  ‘Voluta’ corrugata Binkhorst 1861 PH90
  ‘Coralliophila’ decussata de Loriol 1882 PH90
  ‘Tritonium’ indicum Spengler 1923 PH90
  ‘Fasciolaria’ lloydi Stanton 1920 PH90
  Petersia medicinensis Cragin 1894 PH90
  ‘Voluta’ nitidula Müller 1851 [incl. Rissoa sandbergeri Müller 1851] PH90
  ‘Mitra’ pristina Yokoyama 1923 PH90
  ‘Buccinella’ quadrata Perry 1811 (n. d.) PH90
  ‘Volutilithes’ quercollis Harris 1896 PH90
  ‘Tritonium’ robustum Geinitz 1874 PH90

Nomina nuda: Admete praecursoria Suter 1915 PH90
             Ovilia ovilia Bast. in Jousseaume 1887 PH90

Admete laevior Leche 1878 PH90 [=A. viridula var. laevior PH90, A. couthouyi laevior O27a; incl. A. middendorffiana Dall 1885 PH90]

Admete minuta [incl. A. minuta var. cancellangulosa Sacco 1894, A. minuta var. expusilla Sacco 1894, A. minuta var. parvillima Sacco 1894, A. minuta var. umbilicatior Sacco 1894] PH90

Admete nassiformis [incl. A. nassiformis var. caudatior Sacco 1894, A. nassiformis var. laevicolumella Sacco 1894, A. nassiformis var. quatuorcostata Sacco 1894, A. nassiformis var. spinosa Sangiorgi 1926] PH90

Admete unalashkensis (Dall 1873) PH90 [=Cancellaria unalashkensis PH90, A. unalaschkensis (l. c.) PH90, C. (Sveltia) unalaskensis O27a]

Admete viridula (Fabricius 1780) [=Tritonium viridulum; incl. A. abnormis Morch & Pulson in Harmer 1918 (n. n.), A. borealis Adams 1855, Cancellaria buccinoides Couthouy 1838 non Sowerby 1832, A. contabulata Friele 1879, Murex costellifer Sowerby 1818, A. costellifera, A. couthonyi (l. c.), Cancellaria couthouyi Jay 1839, A. viridula var. couthouyi, A. costellifera var. crassicosticillata Sacco 1894, *A. crispa Möller 1842, A. crispata (l. c.), A. viridula var. distincta Leche 1878, A. viridula var. elongata Leche 1878, A. costellifera var. fusoconvexa Sacco 1894, A. viridula var. grandis Mörch 1869, A. viridula var. producta Sars 1878, A. viridula var. undata Leche 1878, A. undatocostata Verkrüzen 1875, A. viridula var. ventricosa Friele 1879] PH90

*Admetula evulsa (Solander 1766) [=Buccinum evulsum, Cancellaria evulsa; incl. Fusus biplicatus Lamarck 1803, Tritonium bruecknerii Boll 1846, Cancellaria buccinea (l. c.), C. evulsa var. buccinovula Sacco 1894, C. buccinula Lamarck 1822, C. evulsa var. cancellatina Sacco 1894, C. evulsa var. costatonodosa Sacco 1894, C. evulsa var. crassevaricosa Sacco 1894, C. evulsa var. exbellardi Sacco 1894, C. evulsa var. minor Beyrich 1856 nec C. dufourii var. minor Grateloup 1847 nec C. hirta var. minor Bronn 1848, C. evulsa var. miolonga Sacco 1894, C. buccinula var. multicostata Bellardi 1841 nec C. uniangulata var. multicostata Bellardi 1841 nec C. nodulosa var. multicostata Bellardi 1841, C. evulsa var. nodosissima Sacco 1894, C. evulsa var. oligocancellata Sacco 1894, C. oligoevulsa Sacco 1894, C. evulsa var. parisiensis Cossmann 1889, C. evulsa var. postera Beyrich 1856, C. posteror (l. c.), Muricites pyrastriformis Schlotheim 1820, C. evulsa var. rotundulatior Sacco 1894, C. evulsa var. taurelegans Sacco 1894, C. evulsa var. tauroelegans, C. evulsa var. taurinia Bellardi 1841 nec C. varricosa var. taurinia Bellardi 1841 nec C. lyrata var. taurinia Bellardi 1841 nec C. uniangulata var. taurinia Bellardi 1841 nec C. nodulosa var. taurinia Bellardi 1841 nec C. bonelli var. taurinia Bellardi 1841, C. evulsa var. turritovula Sacco 1894, C. evulsa var. vera Beyrich 1856] PH90

*Aneurystoma dufourii (Grateloup 1832) [=Cancellaria dufourii; incl. C. dufourii var. major Grateloup 1847, C. dufourii var. minor Grateloup 1847] PH90

Aphera bronni [incl. A. bronni var. afenestrata Sacco 1894, A. bronni var. subareticulata Sacco 1894] PH90

Babylonella pusilla (Philippi 1843) [=Fasciolaria pusilla; incl. C. elegans Karsten 1849 nec Sowerby 1822 nec Deshayes 1835, C. elongata (l. c.) nec Nyst 1845 nec C. geslini var. elongata Grateloup 1847, Fusus exilis Philippi 1843 non Menke 1843] PH90

Bivetia martini Cossmann 1899 [=Triton buccinoides Martin 1879 non Cancellaria buccinoides Sowerby 1832, Cancellaria neglecta Martin 1895 non Michelotti 1861] PH90

*Bivetiella similis (Sowerby 1833) [=Cancellaria similis, *Bivetia similis, C. assimile (l. c.), C. assimilis (l. c.), C. similaris (l. c.); incl. C. similis alba Sowerby in Domenech, Espinosa et al. 1983 (n. n.) non Webb 1936, Bivetia bivet Jousseaume 1887] PH90

*Bivetopsia chrysostoma (Sowerby 1832) [=Cancellaria chrysostoma; incl. C. multiplicata Lesson 1841 non Lea 1833] PH90

Brocchinia marshalli (Vignal in Cossmann 1921) PH90 [=Cerittudea (l. c. for Cerithidea) marshalli PH90, Zebittium marshalli F27a, Cerithidea minuta Marshall 1919 non Gabb 1873 PH90]

Brocchinia parvula tauroparva Sacco 1894 [incl. B. mitraeformis var. anodosomagna Sacco 1894, B. mitraeformis var. costata Harmer 1918, B. mitraeformis var. depresseplicata Sacco 1894, B. mitraeformis var. laevifasciata Sacco 1894, Voluta mitraeformis Brocchi 1814 non Lamarck 1811, *B. mitraeformis, B. mitraeformis var. paucicostulata Sacco 1894, B. mitraeformis var. pyramidalis Sacco 1894, B. mitraeformis var. subanodosa Sacco 1894, B. mitraeformis var. subasuturata Sacco 1894] PH90

‘Cancellaria’ indentata Sowerby 1832 [=C. unidentata (l. c.); incl. C. affinis Reeve 1856 non Adams 1852, *Bivetia mariei Jousseaume 1887, B. mariae (l. c.)] PH90

‘Cancellaria’ laticostata Tenison-Woods 1880 [incl. C. platypleura Tate 1898, *Gergovia platypleura] PH90

‘Cancellaria’ undulata Sowerby 1849 [incl. *Sydaphera renovata Iredale 1929, C. undulata var. truncata Sowerby 1849] PH90

*Contortia contorta (Basterot 1825) [=Cancellaria contorta; incl. Ca. contorta var. altavillae Libassi 1859 nec C. altavillae Aradas 1846 nec C. labrosa var. altavillae Libassi 1859, Ca. contorta var. dertocacellensis Sacco 1894, Ca. contorta var. dertofusula Sacco 1894, Ca. contorta var. fusosimplex Sacco 1894, Ca. contorta var. ligeriana Glibert 1952, Ca. contorta var. taurinia Bellardi 1841 nec C. varricosa var. taurinia Bellardi 1841 nec C. lyrata var. taurinia Bellardi 1841 nec C. uniangulata var. taurinia Bellardi 1841 nec C. nodulosa var. taurinia Bellardi 1841 nec C. bonelli var. taurinia Bellardi 1841 nec C. evulsa var. taurinia Bellardi 1841 nec C. cancellata var. taurinia Bellardi 1841, Ca. contorta var. tauroturritula Sacco 1894] PH90

*Coptostoma quadratum (Sowerby 1822) [=Cancellaria quadrata; incl. Co. quadratum f. carinatum Wrigley 1935, Ca. quadrata var. planistria von Koenen 1889 non C. planistria von Koenen 1885, Co. quadratum var. porrectum Wrigley 1935] PH90

*Crawfordina crawfordiana (Dall 1891) [=Cancellaria crawfordiana, *Crawfordia crawfordiana; incl. Ca. crawfordiana var. fugleri Arnold 1907, Ca. oregonesis Dall 1909 non Conrad 1865] PH90

*Gulia acutangula (Faujas de Saint Fond 1817) [=Cancellaria acutangula; incl. C. acutangularis Lamarck 1822, C. acutangula var. depressicosta Sacco 1894, C. acutangula var. longojuvenis Sacco 1894, C. acutangulata var. polonica Pusch 1837, Mitra scalariformis Borson 1825] PH90

*Hertleinia mitriformis (Sowerby 1832) [=Cancellaria mitriformis, C. mitraeformis (l. c.) non Buccinum mitraeforme Pusch in Andrzejowski 1830, C. sowerbyi Crosse 1861; incl. C. uniplicata Sowerby 1832] PH90

*Inermia inermis (Pusch 1837) [=Cancellaria inermis, Buccinum mitraeforme Pusch in Andrzejow 1830 non Voluta mitraeformis Brocchi 1814, C. mitraeformis; incl. C. inermis var. angulata Sieber 1936 non C. angulata Eichwald 1830, C. inermis var. colligens Sacco 1894] PH90

*Mariasveltia lunata (Conrad 1830) [=Cancellaria lunata; incl. C. scalarina Conrad 1866 nec Lamarck 1822 nec Sowerby 1849] PH90

*Misteia planispira (Nyst 1845) [=Cancellaria planispira, C. planospira (l. c.) non C. obesa var. planospira Grant & Gale 1931] PH90

*Narona clavatula (Sowerby 1832) [=Cancellaria clavatula, C. calvatula (l. c.), *Panarona clavatula; incl. Narona hidalgoi Jousseaume 1887] PH90

*Ovilia doliolaris (Basterot 1825) [=Cancellaria doliolaris, O. doliaris (l. c.); incl. C. doliolaris var. umbilicina Sacco 1894] PH90

*Plesiocerithium maglorii (Melleville 1843) [=Cancellaria maglorii, P. magloirei (l. c.); incl. Turbinella ornata Watelet 1853] PH90

Plesiotriton deshayesianus Beu & Maxwell 1987 [=Triton turriculatum Deshayes 1835 non Deshayes 1833; incl. T. turriculatum var. latum Dareste de la Chavanne 1910] PH90

Scalptia hystrix (Reeve 1856) [=Cancellaria hystrix, S. histrio (l. c.); incl. S. macconkeyi Jousseaume 1894] PH90

*Scalptia obliquata (Lamarck 1822) [=Cancellaria obliquata, C. obliqua (l. c.); incl. C. asperula Deshayes 1830] PH90

Scalptia scala (Gmelin 1791) [=Murex scala; incl. Cancellaria costata Sowerby 1833 non Sowerby 1822, Trigonaphera withrowi Petit 1976] PH90

*Type species of generic name indicated

REFERENCES

[BR05] Bouchet, P., & J.-P. Rocroi. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1–2): 1–397.

[BC01] Boyko, C. B., & J. R. Cordeiro. 2001. Catalog of Recent type specimens in the Division of Invertebrate Zoology, American Museum of Natural History. V. Mollusca, part 2 (class Gastropoda [exclusive of Opisthobranchia and Pulmonata], with supplements to Gastropoda [Opisthobranchia], and Bivalvia). Bulletin of the American Museum of Natural History 262: 1–170.

[F27a] Finlay, H. J. 1927a. A further commentary on New Zealand molluscan systematics. Transactions and Proceedings of the New Zealand Institute 57: 320–485.

[F27b] Finlay, H. J. 1927b. New specific names for austral Mollusca. Transactions and Proceedings of the New Zealand Institute 57: 488–533.

[O27a] Oldroyd, I. S. 1927a. The Marine Shells of the West Coast of North America vol. II pt I. Stanford University Press: Stanford University (California).

[O27b] Oldroyd, I. S. 1927b. The Marine Shells of the West Coast of North America vol. II pt. III. Stanford University Press: Stanford University (California).

[POH88] Pawlik, J. R., J. B. O’Sullivan & M. G. Harasewych. 1988. The egg capsules, embryos, and larvae of Cancellaria cooperi (Gastropoda: Cancellariidae). Nautilus 102 (2): 47–53.

[PH90] Petit, R. E., & M. G. Harasewych. 1990. Catalogue of the superfamily Cancellarioidea Forbes and Hanley, 1851 (Gastropoda: Prosobranchia). Nautilus Supplement 1: 1–69.

[S11] Simone, L. R. L. 2011. Phylogeny of the Caenogastropoda (Mollusca), based on comparative morphology. Arquivos de Zoologia 42 (4): 161–323.

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