Parachipteria punctata, copyright R. Penttinen.

Belongs within: Circumdehiscentiae.
Contains: Achipteria, Campachipteria, Anachipteria.

The Achipterioidea is a group of oribatid mites known from soil, litter, mosses and liverworts. Members of the Achipteriidae have characteristic spiral internal thickenings in the bothridium and a distinct long, thin, dorsally concave genu in leg IV (Norton & Behan-Pelletier 2009). Members of the Tegoribatidae have a fused lamella, and pteromorphs that are hinged and movable (Behan-Pelletier 2001). The Epactozetidae are a Neotropical family with large, auriculate pteromorphs and foveate patterning on the notogaster (Norton & Behan-Pelletier 2009).

Characters (from Norton & Behan-Pelletier 2009): Prodorsum with lamellae, adjacent medially or fused medially or separated; lamellar cusps, tutorium, translamella present or absent. Genal notch present or absent. Dorsophragmata and pleurophragmata present. Chelicerae chelate-dentate; palpal eupathidium acm attachned to solenidion; subcapitular mentum with or without complete tectum. Axillary saccule of subcapitulum usually absent. Discidium and custodium present, postanal porose area present or absent. Pedotectum I large, extending to base of bothridium; pedotectum II present. Notogaster with or without knifelike humeral process; with tectum posteriorly, overlapping medially or not; pteromorphs present, with or without hinge; lenticulus present. Notogastral setation ten pairs. Octotaxic system developed as porose areas or saccules, sometimes minute. Genital setation six pairs. Bothridium with or without internal, spiral thickenings. Genu IV often concave dorsally, subequal in length or longer than tibia IV; tibia IV with or without solenidion, with or without carina on dorsal surface. Immatures plicate, apheredermous or eupheredermous; integument at least partially tuberculate or not; unideficient. Humeral organ absent. Integument surrounding opening of opisthonotal gland porose or not. Seta d present to tritonymph on tibiae and genua I-IV, or only IV, when respective solenidion exists. Protonymphal legs IV setation 0-0-0-0-7, 0-1-0-0-7, or 0-0-2-0-7. Retrotecta on tibiae and tarsi present or absent.

    |--Epactozetidae NB-P09
    |    |--Epactozetes Grandjean 1930 BB92
    |    |    |--*E. imitator Grandjean 1930 BB92
    |    |    `--E. setosus Balogh & Mahunka 1969 S04
    |    `--Truncozetes Balogh & Mahunka 1969 BB92
    |         |--*T. mucronatus Balogh & Mahunka 1969 BB92
    |         |--T. rugosus Mahunka 1998 S04
    |         `--T. sturmi Balogh 1984 S04
    |--Achipteriidae B-P01
    |    |--Achipteria S04
    |    |--Campachipteria S04
    |    |--Anachipteria S04
    |    |--Hoffmanacarus Mahunka 1995 S04
    |    |    `--*H. virginianus Mahunka 1995 S04
    |    |--Cubachipteria Balogh & Mahunka 1979 S04
    |    |    |--*C. remota (Balogh & Mahunka 1979) [=Achipteria (*Cubachipteria) remota] S04
    |    |    `--C. bispina (Calugar 1990) [=Achipteria (Cubachipteria) bispina] S04
    |    |--Dentachipteria Nevin 1974 S04
    |    |    |--*D. ringwoodensis Newin 1974 S04
    |    |    `--D. highlandensis Nevin 1974 S04
    |    |--Novoribatella Engelbrecht 1986 NB-P09, S04
    |    |    |--*N. transcripta (Mahunka 1985) [=Tectoribates transcriptus] S04
    |    |    |--N. minutisetarum Engelbrecht 1986 S04
    |    |    `--N. pentasetarum Engelbrecht 1986 S04
    |    |--Tectoribates Berlese 1910 NB-P09, S04 [incl. Anoribatella Kunst 1962 S04]
    |    |    |--*T. proximus (Berlese 1910) (see below for synonymy) S04
    |    |    |--T. africanus Balogh 1959 S04
    |    |    |--T. deserticola (Balogh & Mahunka 1965) [=Anoribatella deserticola] S04
    |    |    |--T. mongolicus Bayartogtokh 1997 S04
    |    |    `--T. ornatus (Schuster 1958) [=Anachipteria ornata] S04
    |    |--Cerachipteria Grandjean 1935 [incl. Zygachipteria Mihelčič 1956] S04
    |    |    |--*C. digita Grandjean 1935 S04
    |    |    |    |--C. d. digita S04
    |    |    |    |--C. d. jugata Mihelčič 1956 [=C. (Zygachipteria) jugata] S04
    |    |    |    `--C. d. pyrenaica Pérez-Íñigo 1991 [incl. C. fusiger Mihelčič 1957 (n. d.)] S04
    |    |    |--C. franzi Willmann 1953 S04
    |    |    `--C. minuscula (Berlese 1902) [=Oribata minuscula] S04
    |    `--Parachipteria Hammen 1952 [incl. Pseudachipteria Travé 1960] S04
    |         |--*P. punctata (Nicolet 1855) [=Oribata punctata] S04
    |         |--P. agenjoi (Pérez-Íñigo 1976) [=Pseudachipteria agenjoi] S04
    |         |--P. floresiana (Pérez-Íñigo 1992) [=Pseudachipteria floresiana] S04
    |         |--P. insularis (Pérez-Íñigo 1992) [=Pseudachipteria insularis] S04
    |         |--P. magna (Sellnick 1928) S04 [=Notaspis magna S04, *Pseudachipteria magna BB92]
    |         |--P. nicoleti (Berlese 1883) S07
    |         `--P. ovalis (Koch 1835) (n. d.) [=Oribata ovalis] S04
    `--Tegoribatidae B-P01
         |--Williamszetes Hammer 1961 [incl. Williamsia Hammer 1958 non Carus 1890] S04
         |    `--*W. elsosneadensis (Hammer 1958) [=Williamsia elsosneadensis] S04
         |--Physobates Hammer 1962 S04
         |    `--*P. spinipes Hammer 1962 S04
         |--Tutoridens Krivolutsky 1980 BB92
         |    `--*T. kirgiljakhicus Druk 1980 BB92
         |--Lemurobates Mahunka 1997 S04
         |    `--*L. antsiranana Mahunka 1997 S04
         |--Paraphysobates Mahunka 1985 [=Monophysobates Balogh & Balogh 1992] S04
         |    |--*P. abditus Mahunka 1985 S04 [=*Monophysobates abditus BB92]
         |    `--P. natalensis (Engelbrecht 1986) [=Tegoribates natalensis] S04
         |--Scutozetes Hammer 1952 [incl. Neolepidozetes Hammer 1977] S04
         |    |--*S. lanceolatus Hammer 1952 S04
         |    `--S. ovalis (Hammer 1977) S04 [=*Neolepidozetes ovalis BB92]
         |--Neophysobates Luxton 1987 (see below for synonymy) S04
         |    |--*N. monodactylus (Hammer 1966) S04 [=Physobates monodactylus S04, *Paraphysobates monodactylus BB92]
         |    |--N. montanus (Engelbrecht 1986) [=Tegoribates montanus] S04
         |    `--N. nudus (Engelbrecht 1986) [=Tegoribates nudus] S04
         |--Umbellozetes Krivolutsky 1969 S04
         |    |--*U. fuscus Krivolutsky 1969 S04
         |    |--U. ombinatus Wang & Solhoy 1999 S04
         |    |--U. petrovae (Golosova & Karppinen 1984) [=Tutoridens petrovae] S04
         |    `--U. squamaceus Wang & Shen 1999 S04
         |--Tegoribates Ewing 1917 [incl. Lepidoribates Sellnick 1920] S04
         |    |--*T. subniger Ewing 1917 S04
         |    |--T. americanus Hammer 1958 S04
         |    |--T. bryophilus Woolley 1965 S04
         |    `--T. latirostris (Koch 1844) [=Zetes latirostris; incl. Oribata alzanii Coggi 1900] S04
         |--Lepidozetes Berlese 1910 [incl. Onazetes Bugrov 1991] S04
         |    |--*L. singularis Berlese 1910 S04
         |    |--L. conjunctus Schweizer 1922 [incl. L. chernovi Rjabinin 1974] S04
         |    |--L. dashidorzsi Balogh & Mahunka 1965 S04
         |    |--L. latipilosus Hammer 1952 S04
         |    |--L. trifolius (Fujikawa 1972) [=Tegoribates trifolius] S04
         |    `--L. umbellatus (Bugrov 1991) [=Onazetes umbellatus] S04
         `--Hypozetes Balogh 1959 NB-P09, S04
              |--*H. imitator Balogh 1959 BB92
              |--H. aviger Hammer 1982 S04
              |--H. brevisetus Mahunak 1987 S04
              |--H. bulgaricus Jeleva 1962 S04
              |--H. laysanensis Aoki 1964 B-P01
              |--H. rostralis Tseng 1984 S04
              `--H. translamellatus Wallwork 1965 S04
                   |--H. t. translamellatus S04
                   `--H. t. saudicus Bayoumi & Al-Khalifa 1984 S04

Neophysobates Luxton 1987 [=Paraphysobates Luxton 1985 non Mahunka 1985; incl. Pseudophysobates Fujikawa 1991] S04

*Tectoribates proximus (Berlese 1910) [=Sphaerozetes (*Tectoribates) proximus; incl. T. punctatus Sellnick 1943] S04

*Type species of generic name indicated


[BB92] Balogh, J., & P. Balogh. 1992. The Oribatid Mites Genera of the World vol. 1. Hungarian Natural History Museum: Budapest.

[B-P01] Behan-Pelletier, V. M. 2001. Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae). In: Halliday, R. B., D. E. Walter, H. C. Proctor, R. A. Norton & M. J. Colloff (eds) Acarology: Proceedings of the 10th International Congress pp. 50–57. CSIRO Publishing: Melbourne.

[NB-P09] Norton, R. A., & V. M. Behan-Pelletier. 2009. Suborder Oribatida. In: Krantz, G. W., & D. E. Walter (eds) A Manual of Acarology 3rd ed. pp. 430–564. Texas Tech University Press.

[S07] Schatz, H. 2007. Biogeography of oribatid mites (Acari, Oribatida) from the Cordillera de Talamanca, Costa Rica and Panama. In: Morales-Malacara, J. B., V. M. Behan-Pelletier, E. Ueckermann, T. M. Pérez, E. G. Estrada-Venegas & M. Badii (eds) Acarology XI: Proceedings of the International Congress pp. 151–167. Instituto de Biología and Faculdad de Ciencias, Universidad Nacional Autónoma de México, Sociedad Latinoamericana de Acarología: México.

[S04] Subías, L. S. 2004. Listado sistemático, sinonímico y biogeográfico de los ácaros oribátidos (Acariformes, Oribatida) del mundo (1758–2002). Graellsia 60 (número extraordinario): 3–305.

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