Nanopsocus oceanicus, photographed by Diane Young.

Belongs within: Troctomorpha.
Contains: Liposcelis, Embidopsocinae, Tapinella.

The Liposcelidoidea is a group of small booklice and barklice with micropterous or wingless males and often females.

Characters (from Mockford 1993, as Nanopsocetae): Small forms, rarely over 2 mm in length. Wings, when present (females only), without scales, with not over two M branches in forewing, never with IIA. Coxal organ absent or at most represented by slight bulge in cuticle. Ovipositor of three valvulae but v2 and v3 fused together in some forms.

Liposcelidoidea [Liposceletae, Liposceloidea, Nanopsocetae] R02
    |--Liposcelidae R02
    |    |  i. s.: Belaphopsocus Badonnel 1955 B55
    |    |           |--*B. vilhenai Badonnel 1955 B55
    |    |           `--B. badonneli B86
    |    |         Belapha [incl. Semnopsocus] B55
    |    |           `--B. schoutedeni Enderlein 1917 B55
    |    |         Troctulus Badonnel 1955 B55
    |    |           `--*T. machadoi Badonnel 1955 B55
    |    |--Liposcelis M93
    |    `--Embidopsocinae M93
    |--Troctopsocoidea L02
    |    |--Coleotroctellus Lienhard 1988 [incl. Sinitroctpsocus Li 1993; Troctopsocidae] L02
    |    |    |--*C. burckhardi Lienhard 1988 L02
    |    |    `--C. sui (Li 1993) [=Sinitroctpsocus sui] L02
    |    `--Sphaeropsocidae R02
    |         |--Sphaeropsocus Hagen 1882 [incl. Paleotroctes Enderlein 1911] M93
    |         |    `--*S. kunowii M93
    |         `--Sphaeropsocopsis Badonnel 1963 M93
    |              |--*S. chilensis M93
    |              |--S. argentinus (Badonnel 1962) [=Sphaeropsocus argentinus] M93
    |              `--S. microps M93
    `--Pachytroctidae [Pachytroctoidea] R02
         |--*Psyllotroctes plaumanni [=Pachytroctes plaumanni] M93
         |--Antilopsocus bilimbatus (Enderlein 1908) L02
         |--Tapinella M93
         |--Peritroctes B55
         |    |--P. angolensis Badonnel 1955 B55
         |    `--P. natalensis B55
         |--Nanopsocus Pearman 1928 [incl. Onychotroctes Badonnel 1969] M93
         |    |--*N. oceanicus Pearman 1928 (see below for synonymy) M93
         |    |--N. falsus M93
         |    |--N. longicornis M93
         |    `--N. trifasciatus M93
         `--Pachytroctes Enderlein 1905 M93
              |--*P. aegyptius Enderlein 1905 M93
              |--P. achrosta S95
              |--P. aglyphus Badonnel 1955 B55
              |--P. ambiguus Badonnel 1955 B55
              |--P. aurantiacus Badonnel 1949 B49
              |--P. australis Ribaga 1911 B49
              |--P. bicoloripes Badonnel 1949 B49
              |--P. brunneus Ribaga 1911 B49
              |--P. dichromoscelis Badonnel 1949 S95
              |--P. dundoensis Badonnel 1955 B55
              |--P. ealensis Badonnel 1949 B55
              |--P. enigmaticus Badonnel 1955 B55
              |--P. granulosus Badonnel 1955 B55
              |--P. infuscatus S95
              |--P. insularis S95
              |--P. ixtapaensis S95
              |--P. maculosus M93
              |--P. natalensis B55
              |--P. neoleonensis García Aldrete 1986 M93
              |--P. nivecinctus Badonnel 1955 B55
              |--P. pacificus M93
              |--P. rugosus S95
              |--P. sericeus S95
              |--P. sinuosus Smithers 1995 S95
              |--P. tapinelloides Badonnel 1955 B55
              `--P. velutina S95

*Nanopsocus oceanicus Pearman 1928 [=Tapinella oceanica; incl. T. africana Badonnel 1948, Onychotroctes africanus, T. pallida Badonnel 1949] M93

*Type species of generic name indicated


[B49] Badonnel, A. 1949. Psocoptères de la Cote d'Ivoire. Mission Paulian-Delamare (1945). Revue Française d'Entomologie 16: 20-46.

[B55] Badonnel, A. 1955. Psocoptères de l'Angola. Diamang Publicações Culturais 26: 11-267.

[B86] Badonnel, A. 1986. Psocoptères de Colombie (Insecta, Psocoptera). Spixiana 9 (2): 179-223.

[L02] Li F. 2002. Psocoptera of China vol. 1. Science Press: Beijing.

[M93] Mockford, E. L. 1993. North American Psocoptera (Insecta). Sandhill Crane Press, Inc.

[R02] Rasnitsyn, A. P. 2002. Superorder Psocidea Leach, 1815. In History of Insects (A. P. Rasnitsyn & D. L. J. Quicke, eds) pp. 125-133. Kluwer Academic Publishers: Dordrecht.

[S95] Smithers, C. N. 1995. Psocoptera (Insecta) of Christmas Island. Invertebrate Taxonomy 9: 529-561.

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