Belongs within: Geometroidea.
The Uraniidae are a somewhat disparate group of moths united by the presence of sexually dimorphic tympanal organs on the abdomen.
Characters (from Nielsen & Common 1991): Small to large; head smooth scaled; ocelli absent; chaetosemata present, large; antennae thickened, dentate or pectinate in male, filiform in female; proboscis present, unscaled; maxillary palps minute, one-segmented; labial palps small, slender, upturned; epiphysis present; spurs 0-2-3 or 0-2-4; fore wing without wing-locking microtrichia, R5 well separated from other branches of R and usually forked with M1; CuP absent; 1A+2A sometimes with minute fork; hind wing with or without frenulum, with or without tails, M2 widely separate from M3, CuP absent, one or two anal veins; abdomen with dimorphic paired tympanal organs situated laterally between S2 and S3 in male and at base of S2 in female. Eggs of upright type. Larva with prolegs on segments 3-6 and 10, secondary setae few or absent, prothorax with two L setae, setae on raised pinacula, spiracles on prothorax and abdominal segment 8 much the largest; external feeder. Pupa with dense cover of fine punctures on abdomen, segments 5 and 6 movable, without dorsal spines, with pointed cremaster and divergent anal spines; in cocoon; not protruded at ecdysis.
| i. s.: Acropteris nanula P27
| Lobogethes interrupta P27
| Alcidis DS73
| |--A. agathyrsus DS73
| `--A. zodiaca DS73
| Nyctalemon DS73
| |--N. orontes P27
| `--N. patroclus DS73
| |--N. p. patroclus DS73
| `--N. p. macleayi [incl. N. p. goldiei] DS73
| Urania DS73
| |--U. croesus DS73
| |--U. fulgens DS73
| |--U. leilus DS73
| |--U. ripheus [incl. Chrysiridia madagascariensis] DS73
| `--U. sloanus DS73
`--Epipleminae [Epiplemidae] GE05
*Type species of generic name indicated
[DS73] Dickens, M., & E. Storey. 1973. The World of Moths. Macmillan Publishing Co., Inc.: New York.
[GE05] Grimaldi, D., & M. S. Engel. 2005. Evolution of the Insects. Cambridge University Press: New York.
Nielsen, E. S., & I. F. B. Common. 1991. Lepidoptera. In: CSIRO. The Insects of Australia, 2nd ed., vol. 2, pp. 817-915. Melbourne University Press.
[P27] Philpott, A. 1927. The maxillae in the Lepidoptera. Transactions and Proceedings of the New Zealand Institute 57: 721-746.