Eupterotidae

Jana tantalus, photographed by Mike Nyenes.


Belongs within: Macrolepidoptera.
Contains: Eupterote.

The Eupterotidae are a mostly Old World family of moths, the majority of species of which are nocturnal though a small number are day-flying.

Characters (from Nielsen & Common 1991): Antennae bipectinate to apex in both sexes; proboscis very weak or absent; maxillary palps absent; labial palps short; epiphysis sometimes present, tibial spurs 0-2-2; fore wing usually with retinaculum, areole absent, R2 and R3 usually fused, and R2 or R2+3 arising from R4 beyond its junction with R5, CuP vestigial or absent; hind wing with frenulum usually present, functional, Sc and Rs well separated, usually connected by R1, CuP vestigial or absent, two anal veins. Larva with dense secondary setae, often branched, dorsal verrucae of abdominal segment 1 simialr to those of 2-8, crochets biordinal, shorter series with subapical spur or dentate. Pupa in flimsy cocoon of silk mixed with larval hairs.

<==Eupterotidae
    |--‘Hemijana’ variegata Rothschild 1917 NO08a
    |--Ganisa Walker 1855 NO08a
    |    `--*G. postica Walker 1855 NO08a
    |--Pandala Walker 1855 NO08a
    |    `--*P. dolosa Walker 1855 NO08a
    |--Melanothrix Felder & Felder 1874 NO08a
    |    `--M. nymphaliaria (Walker 1866) [=Gnophos nymphaliaria; incl. *M. pulchricolor Felder & Felder 1874] NO08a
    |--Pseudoganisa Schultze 1910 NO08a
    |    `--*P. currani Schultze 1910 [incl. Apha gonioptera (West 1932)] NO08a
    |--Pseudojana Hampson 1893 NO08a
    |    |--*P. incandescens (Walker 1855) [=Jana incandescens] NO08a
    |    `--P. clemensi Schultze 1907 ONE03
    |--Poloma Walker 1855 [incl. Sarvena Walker 1865] NO08a
    |    |--*P. angulata Walker 1855 NO08a (see below for synonymy)
    |    `--P. nahor (Druce 1896) [=Phasicnecus nahor, Stenoglene nahor] NO08a
    |--Preptos Schaus 1892 NO08a
    |    |--*P. oropus Schaus 1892 [incl. P. norax Druce 1893] NO08a
    |    `--P. hidalgoensis Beutelspacher-B. 1983 NO08a
    |--Neopreptos Draudt 1930 NO08a
    |    |--*N. clazomenia (Druce 1886) [=Coloradia clazomenia] NO08a
    |    `--N. marathusa (Druce 1886) [=Coloradia marathusa] NO08a
    |--Apona Walker 1856 [incl. Pachyclea Gistel 1857] NO08a
    |    |--‘Gastropacha’ caschmirensis Kollar in Kollar & Redtenbacher 1844 (see below for synonymy) NO08a
    |    |--A. gyraleus (Orhant 2000) [=Ganisa gyraleus] NO08a
    |    `--A. ligustri (Mell 1930) NO08a
    |--Apha Walker 1855 [incl. Preptothauma Draudt 1931] NO08a
    |    |--*A. subdives Walker 1855 NO08a
    |    |--‘Brachytera’ aequalis Felder & Felder 1874 NO08a
    |    |--A. floralis Butler 1881 NO08a
    |    `--*Preptothauma’ oxydiata Draudt 1931 NO08a
    |--Panacela Walker 1865 (see below for synonymy) NO08a
    |    |--*P. rufescens Walker 1865 NO08a
    |    |--P. lewinae (Lewin 1805) (see below for synonymy) NO08a
    |    |--P. nyctopa (Turner 1922) [=*Mallodeta nyctopa] NO08a
    |    `--P. syntropha Turner 1922 ONE03
    |--Eupterotinae [Cotanini] NO08a
    |    |--Eupterote NO08a
    |    |--Palirisa Moore 1884 NO08a
    |    |    `--*P. lineosa (Walker 1855) [=Jana lineosa] NO08a
    |    |--Dreata Walker 1855 NO08a
    |    |    `--*D. hades Walker 1855 NO08a
    |    |--Rarisquamosa Bethune-Baker 1910 NO08a
    |    |    `--*R. arfaki Bethune-Baker 1910 NO08a
    |    |--Sangatissa Moore 1883 NO08a
    |    |    `--*S. subcurvifera (Walker 1865) NO08b [=Dreata subcurvifera NO08b, Messata subcurvifera NO08a]
    |    |--Nisaga Walker 1855 NO08a
    |    |    |--*N. simplex Walker 1855 NO08a
    |    |    `--N. rufescens Hampson 1895 NO08a
    |    |--Paracydas Bethune-Baker 1908 NO08a
    |    |    |--*P. biagi Bethune-Baker 1908 NO08a
    |    |    `--P. postpallida Rothschild 1917 NO08a
    |    |--Lasiomorpha Joicey & Talbot in Joicey et al. 1916 NO08a
    |    |    |--*L. noakesi Joicey & Talbot in Joicey et al. 1916 NO08a
    |    |    `--L. meeki (Rothschild 1917) NO08a
    |    |--Melanergon Bethune-Bake 1904 NO08a
    |    |    `--M. vidua (Walker 1865) [=Cotana vidua; incl. *M. proserpina Bethune-Baker 1904] NO08a
    |    |         |--M. v. vidua NO08a
    |    |         `--M. v. fergussonis Rothschild 1917 NO08a
    |    |--Cotana Walker 1865 (see below for synonymy) NO08a
    |    |    |--*C. rubrescens Walker 1865 [incl. *Nervicompressa unistrigata Bethune-Baker 1904] NO08a
    |    |    |--*Hypercydas’ calliloma Turner 1903 NO08a
    |    |    |--‘Nervicompressa’ albomaculata Bethune-Baker 1904 [incl. *Epicydas ovata Bethune-Baker 1908] NO08a
    |    |    `--C. serranotata (Lucas 1894) ONE03
    |    `--Sphingognatha Felder & Felder 1874 NO08a
    |         |--*S. asclepiades Felder & Felder 1874 NO08a
    |         |--‘Pseudojana’ pallidipennis Hampson 1895 NO08a
    |         |    |--P. p. pallidipennis NO08a
    |         |    `--P. p. grandis Rothschild 1917 NO08a
    |         `--S. strix (Bryk 1944) [=Apha strix] NO08a
    |--Janinae [Janidae] NO08a
    |    |--Tissanga Aurivillius in Distant 1903 [Tissanginae, Tissangini] NO08a
    |    |    `--*T. pretoriae (Distant 1892) [=Sangatissa pretoriae] NO08a
    |    |--Hemijana Aurivillius 1901 NO08a
    |    |    `--*H. subrosea (Aurivillius 1893) [=Jana subrosea] NO08a
    |    |--Camerunia Aurivillius 1893 NO08a
    |    |    `--‘Homochroa’ orphne Schaus in Schaus & Clements 1893 [incl. *Camerunia insignis Aurivillius 1893] NO08a
    |    |--Epijana Holland 1893 NO08a
    |    |    `--E. cosima (Plotz 1880) [incl. *E. lanosa Holland 1893] NO08a
    |    |--Parajana Aurivillius 1906 NO08a
    |    |    `--*P. gabunica (Aurivillius 1892) (see below for synonymy) NO08a
    |    |--Pterocerota Hampson 1905 NO08a
    |    |    `--*P. virginea Hampson 1905 NO08a
    |    |--Drepanojana Aurivillius 1893 NO08a
    |    |    `--*D. fasciata Aurivillius 1893 NO08a
    |    |--Vianga Fletcher 1982 [=Viana Walker in Chapman 1869 non Adams & Adams 1856] NO08a
    |    |    `--*V. velutina (Walker in Chapman 1869) [=*Viana velutina] NO08a
    |    |--Catajana Strand 1910 NO08a
    |    |    `--*C. bimaculata (Dewitz 1879) [=Dreata bimaculata, Hemijana bimaculata] NO08a
    |    |--Urojana Gaede 1915 NO08a
    |    |    `--*U. eborea Gaede 1915 NO08a
    |    |--Gonojana Aurivillius 1893 NO08a
    |    |    `--*G. dimidiata Aurivillius 1893 NO08a
    |    |--Hibrildes Druce 1888 [incl. Anengya Karsch 1895; Hibrildidae, Hibrildinae, Hibrildini] NO08a
    |    |    |--*H. norax Druce 1888 [incl. *Anengya spiritalis Karsch 1895] NO08a
    |    |    `--H. crawshayi Butler 1896 [incl. H. ansorgei Kirby 1896, H. venosus Kirby 1896] NO08a
    |    |--Acrojana Aurivillius 1901 NO08a
    |    |    |--*A. sciron (Druce 1888) [=Jana sciron] NO08a
    |    |    `--A. rosacea (Butler 1874) NO08a
    |    |--Hoplojana Aurivillius 1901 NO08a
    |    |    |--*H. indecisa Aurivillius 1901 NO08a
    |    |    |--H. rhodoptera (Gerstäcker 1871) ONE03
    |    |    `--H. zernyi Gschwandner 1923 NO08a
    |    |--Stenoglene Felder & Felder 1874 [incl. Phasicnecus Butler 1894] NO08a
    |    |    |--*S. hilaris Felder & Felder 1874 NO08a
    |    |    |--*Phasicnecus’ gregorii Butler 1894 NO08a
    |    |    |--S. obtusa (Walker 1865) ONE03 [incl. S. tristis Felder & Felder 1874 NS07]
    |    |    `--S. rosea (Druce 1886) ONE03
    |    `--Jana Herrich-Schäffer 1854 NO08a
    |         |--*J. eurymas Herrich-Schäffer 1854 NO08a
    |         |--J. caesarea Weymer 1909 NO08a
    |         |--J. palliatella Viete 1954 NO08a
    |         |--J. tantalus Herrich-Schäffer 1854 ONE03
    |         |--J. tripunctata Aurivillius 1897 [=Hoplojana tripunctata] NO08a
    |         |    |--J. t. tripunctata NO08a
    |         |    `--J. t. roseata Rothschild 1917 NO08a
    |         `--J. yokoana (Bethune-Baker 1927) [=Dreata yokoana] NO08a
    `--Striphnopteryginae [Phialidae, Phialini, Striphnopteryges, Striphnopterygidae] NO08a
         |--Ebbepterote Oberprieler, Nässig & Edwards 2003 NO08a
         |    `--*E. expansa (Lucas 1891) NO08a (see below for synonymy)
         |--Striphnopteryx Wallengren 1858 NO08a
         |    `--*S. edulis (Boisduval 1847) [=Bombyx edulis] NO08a
         |--Trichophiala Aurivillius 1879 NO08a
         |    `--*T. devylderi Aurivillius 1879 NO08a
         |--Marmaroplegma Wallengren 1860 NO08a
         |    `--*M. paragarda Wallengren 1860 [=M. paraguda (l. c.), Rhabdosia paragarda] NO08a
         |--Cyrtojana Aurivillius 1911 NO08a
         |    `--*C. trilineata Aurivillius 1911 NO08a
         |--Schistissa Aurivillius 1901 NO08a
         |    `--*S. uniformis Aurivillius 1901 NO08a
         |--Paraphyllalia Gaede 1927 NO08a
         |    `--*P. degenera (Walker 1855) [=Dreata degenera] NO08a
         |--Bantuana Distant 1906 NO08a
         |    `--*B. cregoei Distant 1906 NO08a
         |--Teratojana Hering 1937 NO08a
         |    `--*T. flavina Hering 1937 NO08a
         |--Rhabdosia Hübner 1820 NO08a
         |    |--*R. vaninia (Stoll in Cramer 1781) [=Phalaena vaninia] NO08a
         |    `--R. patagiata (Aurivillius 1911) ONE03
         |--Lichenopteryx Felder & Felder 1874 NO08a
         |    |--*L. despecta Felder & Felder 1874 NO08a
         |    `--L. scotina Hering 1932 NO08a
         |--Janomima Aurivillius 1901 NO08a
         |    |--*J. mariana (White 1843) [=Bombyx mariana] NO08a
         |    `--J. westwoodi Aurivillius 1902 ONE03
         |--Papuapterote Nässig & Oberprieler 2008 NO08a
         |    |--*P. styx (Bethune-Baker 1908) [=Eupterote styx, Apona styx, Sphingognatha styx, Tagora styx] NO08a
         |    |--P. crenulata (Joicey & Talbot 1916) NO08a
         |    `--P. punctata (Joicey & Talbot 1916) NO08a
         |--Phyllalia Walker 1855 [incl. Homochroa Wallengren 1858 non Hübner 1825] NO08a
         |    |--P. patens (Boisduval 1847) NO08a (see below for synonymy)
         |    |--P. acuta Strand 1911 NO08a
         |    |--P. alboradiata Aurivillius 1911 NO08a
         |    |--P. flavicostata Fawcett 1903 ONE03
         |    |--‘Homochroa’ ornata Felder & Felder 1874 NS07
         |    |--P. thunbergii (Boisduval 1847) ONE03
         |    `--P. valida (Felder & Felder 1874) [=Homochroa valida] NS07
         `--Phiala Wallengren 1860 (see below for synonymy) NO08a
              |--P. costipuncta (Herrich-Schäffer 1855) (see below for synonymy) NO08a
              |--P. arrecta (Distant 1899) ONE03
              |--P. cunina (Cramer 1780) (see below for synonymy) NO08a
              |--P. dasypoda Wallengren 1860 NO08a
              |--‘Euchera’ esomelana Bethune-Baker 1927 NO08a
              |--P. incana (Distant 1897) ONE03
              `--P. sigyna (Aurivillius 1893) [=Jana sigyna, Stenoglene sigyna] NO08a

Cotana Walker 1865 [incl. Epicydas Bethune-Baker 1908, Hypercydas Turner 1903, Nervicompressa Bethune-Baker 1904] NO08a

*Ebbepterote expansa (Lucas 1891) NO08a [=Darala expansa NO08a, Eupterote expansa ONE03; incl. Eu. doddi Turner 1911 ONE03]

‘Gastropacha’ caschmirensis Kollar in Kollar & Redtenbacher 1844 [=*Pachyclea caschmirensis; incl. *Apona pallida Walker 1856] NO08a

Panacela Walker 1865 [incl. Lewinibombyx Strand 1929, Mallodeta Turner 1922 non Butler 1876, Semuta Walker 1865; Panacelinae] NO08a

Panacela lewinae (Lewin 1805) [=Bombyx lewinae, *Lewinibombyx lewinae, Ocinara lewinae; incl. *Semuta pristina Walker 1865] NO08a

*Parajana gabunica (Aurivillius 1892) [=Jana gabunica; incl. P. lamani Aurivillius 1906, J. marmorata Holland 1893] NO08a

Phiala Wallengren 1860 (nom. cons. prop.) [incl. Euchera Hübner 1825 (nom. rej. prop.), Heteromorpha Herrich-Schäffer 1855 nec Hübner 1822 nec Kirby 1823 nec Hodgson 1843, Stibolepis Butler 1878] NO08a

Phiala costipuncta (Herrich-Schäffer 1855) [=*Heteromorpha costipuncta; incl. Dasychira atomaria Walker 1855, *Phiala xanthosoma Wallengren 1860] NO08a

Phiala cunina (Cramer 1780) [=Phalaena cunina, *Euchera cunina, Stibolepis cunina; incl. *S. nivea Butler 1878] NO08a

Phyllalia patens (Boisduval 1847) NO08a [=Bombyx patens NO08a, *Homochroa patens NO08a; incl. Cirina cana Felder & Felder 1874 NS07, Dreata concolor Walker 1855, *Phyllalia concolor Walker 1855 non Dreata concolor Walker 1855 NO08a, Cirina forda (Westwood 1849) NO08a]

*Poloma angulata Walker 1855 NO08a [incl. *Sarvena incompta Walker 1865 NO08a, Homochroa janula Felder & Felder 1874 NSp07]

*Type species of generic name indicated

REFERENCES

[NO08a] Nässig, W. A., & R. G. Oberprieler. 2008a. An annotated catalogue of the genera of Eupterotidae (Insecta, Lepidoptera, Bombycoidea). Senckenbergiana Biologica 88 (1): 53-80.

[NO08b] Nässig, W. A., & R. G. Oberprieler. 2008b. Errata et addenda. Senckenbergiana Biologica 88 (2): 124.

[NS07] Nässig, W. A., & W. Speidel. 2007. On the authorships of the Lepidoptera Atlas of the "Reise der Novara", with a list of the taxa of Bombycoidea [s. l.] therein described (Insecta, Lepidoptera, Bombycoidea). Senckenbergiana Biologica 87 (1): 63-74.

Nielsen, E. S., & I. F. B. Common. 1991. Lepidoptera. In: CSIRO. The Insects of Australia, 2nd ed., vol. 2, pp. 817-915. Melbourne University Press.

[ONE03] Oberprieler, R. G., W. A. Nässig & E. D. Edwards. 2003. Ebbepterote, a new genus for the Australian ‘Eupterote’ expansa (T. P. Lucas), with a revised classification of the family Eupterotidae (Lepidoptera). Invertebrate Systematics 17: 99-110.

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