Achatinoidea

Giant African land snail Achatina fulica, photographed by Alexander R. Jenner.


Belongs within: Stylommatophora.

The Achatinoidea is a group of snails characterised by the absence of a stimulator in the genitalia.

Synapomorphies (from Tillier 1989): Ureteric tube closed; cerebral commisure distinctly shorter than cerebral ganglia width; left parietal ganglion contiguous with visceral ganglion; cerebro-pedal connectives symmetrical; stimulator absent.

<==Achatinoidea [Achatinacea]
    |--Micractaeon Verdcourt 1993 [Micractaeonidae] BR05
    |--Ferussaciidae [Ferrusaciidae, Ferussaciinae] BR05
    |    |--Ferussacia Risso 1826 [=Ferrussacia (l. c.)] BR05
    |    `--Cecilioides Férussac 1814 (see below for synonymy) BR05
    |         |--C. acicula CC06
    |         `--C. eulima CC06
    |--Achatinidae [Achatinae, Achatinida] BR05
    |    |  i. s.: Leptocala mollicella FGN07
    |    |--Callistoplepa Ancey 1888 [=Callistopepla (l. c.); Callistopeplinae, Callistoplepinae] BR05
    |    |--Limicolaria Schumacher 1817 [Limicolariinae] BR05
    |    |    `--L. tenebrica A66a
    |    `--Achatininae BR05
    |         |--Urceus Klein ex Mörch 1857 [Urceidae] BR05
    |         `--Achatina Lamarck 1799 BR05
    |              |--A. balteata Reeve 1849 FGN07
    |              |--A. fulica Bowditch 1822 S88
    |              |--A. leakeyi P02
    |              `--A. turricula Mighels 1845 J49
    `--Subulinidae [Subulinoidea] BR05
         |  i. s.: Oreohomorus P02
         |         Subuliniscus P02
         |         Subulona decollata (Morelet 1873) FGN07
         |         Ischnoglessula FGN07
         |         Striosubulina FGN07
         |         Pseudopeas Putzeys 1899 S88
         |         Curvella FGN07
         |         Nothapalus FGN07
         |         Dictyoglessula FGN07
         |         Oleata FGN07
         |         Pileata FGN07
         |         Kempioconcha FGN07
         |         Lamellaxis S88
         |           |--L. clavulinus (Potiez & Michaud 1838) [=Bulimus clavulinus, Opeas clavulinum] S88
         |           |--L. gracilis (Hutton 1834) S88 (see below for synonymy)
         |           `--L. mauritianus (Pfeiffer 1852) S88
         |         Eremopeas Pilsbry 1906 S88
         |           |--E. interioris (Tate 1894) [=Stenogyra interioris, Pseudopeas (Eremopeas) interioris] S88
         |           `--E. tuckeri (Pfeiffer 1846) (see below for synonymy) S88
         |         Prosopeas achatinaceum (Pfeiffer 1846) (see below for synonymy) S88
         |--Glessula Martens 1860 [Glessulidae, Glessulinae] BR05
         |--Petriola Dall 1905 [Petriolinae] BR05
         |--Rishetia Godwin-Austen 1920 [Rishetiinae] BR05
         |--Subulina Beck 1837 [Subulininae] BR05
         |    `--S. octona (Bruguière 1792) S88 [=Bulimus octonus S88, Rumina (Subulina) octona A66b]
         |--Rumina Risso 1826 [Ruminidae, Rumininae] BR05
         |    `--R. decollata CC06
         |--Opeas Albers 1850 [Opeatinae] BR05
         |    |--O. pumilum (Pfeiffer 1840) (see below for synonymy) S88
         |    `--O. pyrgula Schmacker & Boettger 1891 S88
         |--Coeliaxinae [Caeliaxidae] BR05
         |    |--Coeliaxis Adams & Angas 1865 [=Caeliaxis (l. c.)] BR05
         |    |--Cryptelasmus Pilsbry 1907 [Cryptelasminae] BR05
         |    `--Pyrgina Greef 1882 [Pyrgininae] BR05
         `--Stenogyrinae [Stenogyridae] BR05
              |--Stenogyra Shuttleworth 1854 BR05
              `--Obeliscus Beck 1837 BR05
                   |--O. pusillus (Adams 1866) [=Rumina (Obeliscus) pusilla] A66b
                   `--O. riparius [=Rumina (Obeliscus) riparia] A66b

Cecilioides Férussac 1814 [=Caecilianella Bourguignat 1856, Caecilioides; Caeciliae, Caecilianellea, Cecilioididae] BR05

Eremopeas tuckeri (Pfeiffer 1846) [=Bulimus tuckeri, Prosopeas (Eremopeas) tuckeri; incl. B. walli Cox 1864] S88

Lamellaxis gracilis (Hutton 1834) S88 [=Bulimus gracile S88, L. (Allopeas) gracilis S88, Opeas gracile S88, Stenogyra gracilis TC89; incl. O. ardelio Iredale 1941 S88, O. oparanum (Pfeiffer 1846) S88]

Opeas pumilum (Pfeiffer 1840) [=Bulimus pumilus; incl. Helix goodalli Miller 1822 non Ferussac 1821, Opeas goodalli] S88

Prosopeas achatinaceum (Pfeiffer 1846) [=Bulimus achatinaceus; incl. Achatina javanica Reeve 1849, Opeas javanicum] S88

*Type species of generic name indicated

REFERENCES

[A66a] Adams, H. 1866a. List of the shells collected by Samuel White Baker, Esq., during his recent explorations in central Africa. Proceedings of the Zoological Society of London 1866: 375-376.

[A66b] Adams, H. 1866b. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440-445.

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[CC06] Cameron, R. A. D., L. M. Cook, G. A. Goodfriend & M. B. Seddon. 2006. Fossil land snails of Porto Santo, Madeiran archipelago: change and stasis in Pleistocene to Recent times. Malacologia 49 (1): 25-59.

[FGN07] Fontaine, B., O. Gargominy & E. Neubert. 2007. Land snail diversity of the savanna/forest mosaic in Lopé National Park, Gabon. Malacologia 49 (2): 313-338.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213-231.

[P02] Pickford, M. 2002. Early Miocene grassland ecosystem at Bukwa, Mount Elgon, Uganda. Comptes Rendus Palevol 1: 213-219.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455-604.

[TC89] Tapparone Canefri, T. 1889. Viaggio di Leonardo Fea in Birmania e regioni vicine. XVIII. – Molluschi terrestri e d'acqua dolce. Annali del Museo Civico di Storia Naturale di Genova, Serie 2a 7: 295-359.

Tillier, S. 1989. Comparative morphology, phylogeny and classification of land snails and slugs (Gastropoda: Pulmonata: Stylommatophora). Malacologia 30 (1-2): 1-303.

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