Oahu tree snail Achatinella mustelina, photographed by Brenden Holland.

Belongs within: Stylommatophora.

The Achatinellidae are an exclusively Indo-Pacific group of very small snails (Stanisic et al. 2010).

Characters (from Stanisic et al. 2010): Shells very small, conical to elongate; whorls generally rounded. Sutures moderately indented; umbilicus usually absent. Lip simple, not reflected. Aperture ovate; columella spirally twisted to straight, may have a small lamella; small lamella also usually present on parietal wall. Apertural barriers varying in size during growth of shell, juveniles generally having more prominent barriers compared to adults. Foot short, broad, with very sticky viscous mucus that can be drawn out in strands.

<==Achatinellidae [Achatinellacea, Achatinelloidea]
    |--Auriculella Pfeiffer 1854 [Auriculellidae, Auriculellinae] BR05
    |    `--‘Bulimus’ armatus Mighels 1845 [incl. Auriculella westerlundiana Ancey 1889] J49
    |--Tekoulina Solem 1972 [Tekoulininae] BR05
    |--Tornatellidinae BR05
    |    |--Tornatellides Pilsbry 1910 [Tornatellidini] BR05
    |    `--Tornatellaria Pilsbry 1910 [Tornatellariini] BR05
    |--Achatinellinae BR05
    |    |--Helicteres Beck 1837 [=Helicter Pease 1862; Helicteridae, Helicterinae] BR05
    |    `--Achatinella Swainson 1828 [Achatinellini] BR05
    |         |--A. accineta (Mighels 1845) [=Achatina accineta, Achatinella accincta Gould 1852] J49
    |         |--A. cingula Mighels 1845 J49
    |         |--A. citrina Mighels in Pfeiffer 1845 J49
    |         |--A. decora [incl. A. vestita Mighels 1845] J49
    |         |--A. mustelina Mighels 1845 J49
    |         |--A. nubilosa Mighels 1845 J49
    |         |--A. plicata Mighels in Pfeiffer 1848 J49
    |         |--A. spirizona [incl. A. baetica Mighels in Pfeiffer 1848 (n. n.)] J49
    |         |--A. turritella [incl. A. inornata Mighels 1845] J49
    |         |--A. venusta Mighels 1845 J49
    |         |--A. virgulata [incl. Bulimus insignis Mighels in Reeve 1850 (n. n.)] J49
    |         `--A. viridans Mighels 1845 J49
    |--Tornatellininae [Tornatellinidae] BR05
    |    |--Tornatellina Pfeiffer 1842 [Tornatellinini] BR05
    |    |    `--T. (Leptinaria) lamellata A66
    |    `--Elasmias Pilsbry 1910 [Elasmiatidae, Elasmiatini] BR05
    |         |--E. manilense (Dohrn 1863) [=Tornatellina manilensis] S88
    |         |--E. terrestris (Brazier 1876) (see below for synonymy) S88
    |         `--E. wakefieldiae (Cox 1868) S88
    |--Pacificellinae [Pacificellidae] BR05
    |    |--Lamellidea Pilsbry 1910 [Lamellideinae, Lamellideini] BR05
    |    `--Pacificellini BR05
    |         |--Pacificella Odhner 1921 BR05
    |         `--Tornatellinops Pilsbry & Cooke 1915 [Tornatellinoptini] BR05
    |              |--T. jacksonensis SC07
    |              `--T. pressus SC07
    `--Elasmatininae [Elasmatinidae] BR05
         |--Antonella Cooke & Kondo 1961 [Antonellini] BR05
         |--Tubuaia Cooke & Kondo 1961 [Tubuaiini] BR05
         `--Elasmatinini BR05
              |--Elasmatina Petit de la Saussaye 1843 BR05
              |--Strobilus Anton 1838 BR05
              `--Pitys Mörch 1852 [Pitysinae, Pitysini] BR05

Elasmias terrestris (Brazier 1876) [=Tornatellina terrestris, E. terrestre; incl. E. eucharis Brazier 1876] S88

*Type species of generic name indicated


[A66] Adams, H. 1866. List of land and freshwater shells collected by Mr. E. Bartlett on the Upper Amazons, and on the River Ucayali, eastern Peru, with descriptions of new species. Proceedings of the Zoological Society of London 1866: 440-445.

[BR05] Bouchet, P., J.-P. Rocroi, J. Frýda, B. Hausdorf, W. Ponder, Á. Valdés & A. Warén. 2005. Classification and nomenclator of gastropod families. Malacologia 47 (1-2): 1-397.

[J49] Johnson, R. I. 1949. Jesse Wedgwood Mighels with a bibliography and a catalogue of his species. Occasional Papers on Mollusks 1 (14): 213-231.

[S88] Solem, A. 1988. Non-camaenid land snails of the Kimberley and Northern Territory, Australia. I. Systematics, affinities and ranges. Invertebrate Taxonomy 4: 455-604.

[SC07] Stanisic, J., R. A. D. Cameron, B. M. Pokryszko & J. C. Nekola. 2007. Forest snail faunas from S. E. Queensland and N. E. New South Wales (Australia): patterns of local and regional richness and differentiation. Malacologia 49 (2): 445-462.

Stanisic, J., M. Shea, D. Potter & O. Griffiths. 2010. Australian Land Snails vol. 1. A field guide to eastern Australian species. Bioculture Press: Mauritius.

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