Gelechioidea

Eriogenes cossoides, copyright CSIRO.

Belongs within: Obtectomera.
Contains: Agapalsini, Blastobasinae, Borkhausenia, Gymnobathra, Oecophorinae, Xyloryctinae, Stathmopodinae, Depressaria, Gelechiidae, Cosmopteriginae, Ethmiinae.

The Gelechioidea is a hyperdiverse group of moths, generally characterised by the presence of a densely scaled proboscis. Approximately 16,500 species are known within the clade, with a correspondingly high diversity of lifestyles, with larvae being phytophagous, detritivorous or even parasitic (Grimaldi & Engel 2005). Over half the species of gelechioids are included in the family Oecophoridae, larvae of which mostly tunnel in stems or other plant parts, or inhabit retreats constructed by joining leaves or among detritus. The Elachistidae, grass-miner moths, are mostly leaf or stem miners in Poales. The ecologically diverse Cosmopterigidae are commonly characterised by asymmetrical genitalia in the males. The Coleophoridae are very small moths with two longitudinal patches on spines on each abdominal tergite that are not obscured by scales (Nielsen & Common 1991).

See also: The butterflies get all the glory (Taxon of the Week: Gelechioidea).

Characters (from Nielsen & Common 1991): Ocelli present or absent; chaetosemata absent; scape often with pecten; proboscis with dense, imbricated scales towards base, rarely reduced; maxillary palps small, mostly 4-segmented, folded over base of proboscis, rarely reduced; labial palps usually recurved, apical segment often exceeding vertex, usually tapering, acute; epiphysis usually present; spurs 0-2-4; fore wing with M rarely present in discal cell, chorda vestigial or absent, retinaculum in female often subcubital; hind wing with venation sometimes reduced, CuA rarely with basal pecten of hairs, CuP often absent, female frenulum usually with three bristles; S2 of tineid type; abdomen often with prominent dorsal spines. Larva with 3 L setae on prothorax, crochets in a circle, penellipse, or rarely in two transverse rows, abdominal setae L1 and L2 approximated; concealed feeders, case-bearing, tunnelling in stems or fruits, gall-forming, leaf-ming, joining foliage, or feeding beneath silken shelter. Pupa usually with labial palps and fore femora concealed, without dorsal abdominal spines, usually segments 5-7 movable in male, 5-6 in female; usually in larval shelter, not protruded from shelter or cocoon at ecdysis.

<==Gelechioidea
    |--+--Depressariidae KP19
    |  |    |--Depressaria GE05
    |  |    `--Psilocorsis reflexella KP19
    |  `--+--Gelechiidae KP19
    |     `--+--Autostichinae [Autostichidae] KP19
    |        |    |--Autosticha KP19
    |        |    `--Procometis NC91
    |        |         |--P. bisculcata NC91
    |        |         `--P. hylonoma P27c
    |        `--Cosmopterigidae O94
    |             |  i. s.: Glaphyristis marmarea P27c
    |             |         Trachydora droserodes P27c
    |             |         Euclemensia GE05
    |             |         Morphotica mirifica ZS10
    |             |         Thectophila Meyrick 1927 M27
    |             |           `--*T. acmotypa Meyrick 1927 M27
    |             |--Antequerinae NC91
    |             |--Walshiinae C70
    |             |--Cosmopteriginae NC91
    |             `--Chrysopeleiinae NC91
    |                  |--Chalotis semnostola NC91
    |                  `--Ithome lassula NC91
    `--Elachistidae GE05
         |  i. s.: Cosmiotes NC91
         |           |--C. aphanta C70
         |           |--C. archaeonoma M83
         |           `--C. synethes NC91
         |         Elachista Treitschke 1833 BR17
         |           |--E. archaeonoma P27c
         |           |--E. exaula P27c
         |           |--E. megerleella F92
         |           |--E. ombrodoca P27c
         |           `--E. thallophora P27c
         |         Ptilochares Meyrick 1887 M87
         |           `--*P. trissodesma Meyrick 1887 M87
         |         Castorura Meyrick 1887 M87
         |           `--*C. chrysias Meyrick 1887 M87
         |         Elachistites inclusus RJ93
         |         Anybia P92
         |         Microperittia P92
         |         Palaeoelachista P92
         |         Praemendesia P92
         |--Ethmiinae K07
         `--Stenomatinae [Stenomatidae] E03
              |--Antaeotricha schlaegeri KP19
              |--Agriophara Rosenstock 1885 E03
              |    |--*A. cinerosa E03
              |    |--A. confertella P27c
              |    `--A. platyscia NC91
              `--Eriogenes Meyrick 1925 E03
                   |--*E. mesogypsa Meyrick 1925 E03
                   |--E. cossoides (Butler 1882) [=Phanaca cossoides; incl. E. meyricki Duckworth 1973] E03
                   `--E. nielseni Edwards 2003 E03

Gelechioidea incertae sedis:
  Metachandidae DS73
  Anomologidae DS73
  Pterolonchidae DS73
  Physoptila [Physoptilidae] C74
  Blastodacnidae NC91
  Symmocidae NC91
    |--Symmoca C70
    `--Oegoconiites borisjaki Kuznetsov 1941 P92
  Holcopogonidae NC91
  Mompha WRM02 [Momphidae NC91, Momphinae]
    `--M. cephalonthiella WRM02
  Agonoxena [Agonoxenidae] NC91
    |--A. argaula NC91
    `--A. phoenicia NC91
  Hypertrophidae NC91
    |--Eupselia satrapella NC91
    `--Hypertropha M87
         |--H. chlaenota Meyrick 1887 M87
         `--H. tortriciformis (see below for synonymy) M86
  Batrachedra [Batrachedridae] NC91
    |--B. agaura P27c
    |--B. arenosella NC91
    |--B. microtoma NC91
    |--B. psithyra L27
    `--B. stegodyphobius R13
  Lecithoceridae [Timyridae] NC91
    |--Lecithocera micromela NC91
    `--Crocanthes NC91
         |--C. perigrapta P27c
         |--C. prasinopis NC91
         `--C. sidonia ZS10
  Scythrididae NC91
    |--Scythropites balticella Rebel 1936 P92
    |--Eretmocera NC91
    |    |--E. dioctis NC91
    |    `--E. flavicincta P27c
    `--Scythris NC91
         |--S. epistrota P27c
         |--S. rhabducha NC91
         `--S. scopolella KP19
  Coleophoridae [Eupistidae] GE05
    |  i. s.: Coleophora YS10
    |           |--C. alcyonipennella NC91
    |           |--C. frischella B88
    |           |--C. laricella F62
    |           |--C. lusitanica Baldizzone & Corley 2004 F05
    |           |--C. malivorella G05
    |           |--C. ochroneura C70
    |           |--C. parthenica G05
    |           `--C. serinipennella NC91
    |         Multicoloria YS10
    |           |--M. eucoleos YS10
    |           `--M. singreni YS10
    |         Corythangela NC91
    |--Agapalsini F05
    |--Blastobasinae AH96
    `--Systrophoecini F05
         |--Systrophoeca Falkovitsh 1972 F05
         |--Orghidania F05
         `--Suireia F05
  Oecophoridae [Cryptolechiadae] GE05
    |  i. s.: Tingena DGH93
    |         Borkhausenia P27a
    |         Petalanthes M27
    |         Gymnobathra P27b
    |         Eulechria baryptera P27c
    |         Nymphostola galactina P27c
    |         Ochlogenes advectella P27c
    |         Machaeritis aegrella P27c
    |         Oxythecta acceptella P27c
    |         Coeranica isabella P27c
    |         Saropla philocala P27c
    |         Protomacha chalcaspis P27c
    |         Chezala conjunctella P27c
    |         Lophopepla igniferella P27c
    |         Zonopetala clerota P27c
    |         Callithauma pyrites P27c
    |         Eutorna pabulicola P27c
    |         Thudaca obliquella P27c
    |         Endorsis lactella P27c
    |         Schiffermuelleria orthopanes P27c
    |         Leptocroca meselectra P27c
    |         Euchersadaula lathriopa P27c
    |         Euthictis chloratma P27c
    |         Thamnosara sublitella P27c
    |         Telanepsia AY04
    |         Thyrocopa O94
    |         Opisina arnosella [incl. Nephantis serinopa] YS10
    |         Hoplomorpha abalienella ZS10
    |         Euchaetis ZS10
    |         Habroscopa iriodes ZS10
    |         Corocosma Meyrick 1927 M27
    |           `--*C. memorabilis Meyrick 1927 M27
    |         Agonopterix P04
    |           |--A. alstromeriana P04
    |           |--A. arenella I92
    |           |--A. heracliana RD77
    |           |--A. muricolorella (Busck 1902) [=Depressaria muricolorella] C52
    |           |--A. oregonensis Clarke 1941 C52
    |           `--A. rosaciliella (Busck 1904) [=Depressaria rosaciliella; incl. A. r. echinopanacis] C52
    |         Piloprepes ZS10
    |           |--P. antidoxa ZS10
    |           `--P. lophoptera P27c
    |         Hierodoris H04
    |           |--H. atychioides H04
    |           `--H. frigida P27c
    |         Proteodes M27
    |           |--P. carnifex P27c
    |           `--P. melagrapha Meyrick 1927 M27
    |         Lathicrossa M27
    |           |--L. leucocentra P27c
    |           `--L. prophetica Meyrick 1927 M27
    |         Machimia P27c
    |           |--M. carnea P27c
    |           `--M. zatrephes P27c
    |         Cryptolechia P27c
    |           |--C. liochroa P27c
    |           |--C. radiosella P27c
    |           `--C. tentoriferella RD77
    |         Coesyra P27c
    |           |--C. apora P27c
    |           |--C. dichroella P27c
    |           `--C. paulinella P27c
    |         Atomotricha P27c
    |           |--A. chloronota P27c
    |           |--A. exsomnis P27c
    |           `--A. ommatias L27
    |         Trachypepla M27
    |           |--T. anastrella L27
    |           |--T. aspidephora L27
    |           |--T. conspicuella L27
    |           |--T. contritella P27c
    |           |--T. euryleucota T27
    |           |--T. indolescens Meyrick 1927 M27
    |           `--T. photinella M27
    |         Izatha P27c
    |           |--I. attactella L27
    |           |--I. convulsella L27
    |           |--I. epiphanes P27c
    |           |--I. mira L27
    |           |--I. peroneanella L27
    |           |--I. picarella L27
    |           `--I. planetella L27
    |         Peritornenta circulatella NC91
    |         Pedois NC91
    |         Tonica effractella NC91
    |         Enchocrates glaucopis NC91
    |         Placocosma resumptella [=Oecophora resumptella; incl. P. hephaestea Meyr. 1883] M86
    |         Antidica pilipes (Butl. 1882) [=Latometus pilipes; incl. A. eriophora Meyr. 1883] M86
    |         Gonionota pyrobola Meyrick 1887 M87
    |         Peritorneuta C70
    |         Cormotypa fascialis C70
    |         Sphaerelictis hepialella C70
    |         Borkhausenites bachofeni RJ93
    |         Depressarites P92
    |         Epiborkhausenites P92
    |         Glesseumeyickia P92
    |         Microsymmocites P92
    |         Neoborkhausenites P92
    |         Palaeodepressaria P92
    |         Paraborkhausenites P92
    |         Symmocites P92
    |         Tubuliferola P92
    |--Oecophorinae NC91
    |--Xyloryctinae NC91
    `--Stathmopodinae NC91

Hypertropha tortriciformis [=Heliodes tortriciformis; incl. Orosana desumptana, Anthoecia divitiosa, Hypertropha thesaurella Meyr. 1880] M86

*Type species of generic name indicated

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Last updated: 2 March 2021.