Kousa dogwood Cornus kousa, from here.

Belongs within: Gunneridae.
Contains: Ericales, Gentianidae.

The Asteridae is one of the major clades of flowering plants. Many members of the clade are sympetalous, with the petals fused to varying degrees from the base. The clade is mainly supported by molecular data, but possible synapomorphies include tenuinucellate and unitegmic ovules, sympetaly, and iridoid compounds (Cantino et al. 2007).

<==Asteridae [Hydrangeales, Sympetalae, Theanae]
    |  i. s.: Fouquieria [incl. Idria; Fouquieriaceae, Fouquieriales] T00
    |         Diapensia KJ02 [Diapensiaceae, Diapensiales T00]
    |           |--D. himalaica O88
    |           `--D. lapponica KJ02
    |         Montiniaceae T00
    |           |--Grevea T00
    |           |--Kaliphora T00
    |           `--Montinia T00
    |         Greyia [Greyiaceae] T00
    |         Francoa [Francoaceae] T00
    |--+--Ericales K01
    |  `--Gentianidae K01
    `--Cornales [Cornanae, Cornineae] BL04
         |  i. s.: Tylerianthus T00
         |         Curtisia [Curtisiaceae] T00
         |--+--Alangium [Alangiaceae] K01
         |  |    `--A. salvifolium SR07
         |  |         |--A. s. var. salvifolium SR07
         |  |         `--A. s. var. hexapetalum SR07
         |  `--Cornaceae K01
         |       |--Marlea [incl. Stylidium Loureiro 1790 (nom. rej.?) non Swartz 1805] BR65
         |       |    `--*Stylidium’ chinense (n. d.) BR65
         |       `--Cornus K01
         |            |--C. alternifolia ZB01
         |            |--C. controversa [incl. C. macrophylla] LO98
         |            |--C. florida ZB01
         |            |--C. kousa ZB01
         |            |--C. mas Linnaeus 1753 CD07
         |            |--C. nuttallii ZB01
         |            |--C. officinalis LO98
         |            `--C. sanguinea G98
         `--+--Nyssaceae T00
            |    |--Davidia [Davidioideae] T00
            |    |--Nyssoideae T00
            |    |    |--Camptotheca T00
            |    |    `--Nyssa K01
            |    `--Mastixioideae [Mastixiaceae] T00
            |         |--Diplopanax T00
            |         |--Mastixia T00
            |         |--Retinomastixia glandulosa JD05
            |         `--Eomastixia bilocularis JD05
            `--+--Loasaceae T00
               |    |--Gronovioideae T00
               |    |--Loasa D03 [Loasoideae T00]
               |    |    `--L. volubilis D03
               |    `--Mentzelioideae T00
               |         |--Eucnide K01
               |         `--Mentzelia T00
               `--Hydrangeaceae T00
                    |  i. s.: Dichroa febrifuga D07
                    |--Philadelphoideae T00
                    |    |--Philadelphus K01
                    |    `--Kirengeshoma T00
                    `--+--Fendlera K01
                       `--Hydrangea [Hydrangeoideae] K01
                            |--H. macrophylla MH98
                            |--H. paniculata T07
                            `--H. scandens D03

*Type species of generic name indicated


[BL04] Barkman, T. J., S.-H. Lim, K. M. Salleh & J. Nais. 2004. Mitochondrial DNA sequences reveal the photosynthetic relatives of Rafflesia, the world’s largest flower. Proceedings of the National Academy of Sciences of the USA 101 (3): 787-792.

[BR65] Black, J. M., & E. L. Robertson. 1965. Flora of South Australia. Part IV. Oleaceae-Compositae. W. L. Hawes, Government Printer: Adelaide.

[CD07] Cantino, P. D., J. A. Doyle, S. W. Graham, W. S. Judd, R. G. Olmstead, D. E. Soltis, P. S. Soltis & M. J. Donoghue. 2007. Towards a phylogenetic nomenclature of Tracheophyta. Taxon 56 (3): E1-E44.

[D07] Dash, S. S. 2007. Useful plants of Kabi Sacred Grove, Sikkim. Bulletin of the Botanical Survey of India 49 (1-4): 79-88.

[D03] Dusén, P. 1903. The vegetation of western Patagonia. In Reports of the Princeton University Expeditions to Patagonia, 1896-1899, vol. 8 – Botany (W. B. Scott, ed.) pp. 1-34. The University: Princeton (New Jersey).

[G98] Gruber, J. 1998. Beiträge zur Systematik der Gattung Dicranolasma (Arachnida: Opiliones, Dicranolasmatidae). I. Dicranolasma thracium Staręga und verwandte Formen aus Südosteuropa und Südwestasien. Annalen des Naturhistorischen Museum in Wien. Serie B. Botanik und Zoologie 100: 489-537.

[JD05] Junge, F. W., M. Dolezych, H. Walther, T. Böttger, A. Kühl, L. Kunzmann, P. Morgenstern, T. Steinberg & R. Stange. 2005. Ein Fenster in Landschaft und Vegetation vor 37 Millionen Jahren: Lithologische, sedimentgeochemische und paläobotanische Befunde aus einem Paläoflusssystem des Weißelsterbeckens. Mauritiana 19 (2): 185-273.

[K01] Kårehed, J. 2001. Multiple origin of the tropical forest tree family Icacinaceae. American Journal of Botany 88 (12): 2259-2274.

[KJ02] Kron, K. A., W. S. Judd, P. F. Stevens, D. M. Crayn, A. A. Anderberg, P. A. Gadek, C. J. Quinn & J. L. Luteyn. 2002. Phylogenetic classification of Ericaceae: Molecular and morphological evidence. Botanical Review 68: 335-423.

[LO98] Lack, H. W., & H. Ohba. 1998. Die Xylothek des Chikusai Kato. Willdenowia 28: 263-276.

[MH98] Morikawa, H., A. Higaki, M. Nohno, M. Takahashi, M. Kamada, M. Nakata, G. Toyohara, Y. Okamura, K. Matsui, S. Kitani, K. Fujita, K. Irifune & N. Goshima. 1998. More than a 600-fold variation in nitrogen dioxide assimilation among 217 plant taxa. Plant, Cell and Environment 21: 180-190.

[O88] Ohba, H. 1988. The alpine flora of the Nepal Himalayas: An introductory note. In The Himalayan Plants vol. 1 (H. Ohba & S. B. Malla, eds) The University Museum, University of Tokyo, Bulletin 31: 19-46.

[SR07] Sankar, R. V., K. Ravikumar, N. M. Ganesh Babu & D. K. Ved. 2007. Botany of Anapady MPCA, Palghat district, Kerala with special emphasis on species of conservation concern. Bulletin of the Botanical Survey of India 49 (1-4): 165-172.

[T00] Thorne, R. F. 2000. The classification and geography of the flowering plants: Dicotyledons of the class Angiospermae (subclasses Magnoliidae, Ranunculidae, Caryophyllidae, Dilleniidae, Rosidae, Asteridae, and Lamiidae). The Botanical Review 66: 441-647.

[T07] Tsurusaki, N. 2007. The Chugoku Mountains – a hotspot of geographical differentiation of species. Taxa 22: 3-14.

[ZB01] Zhang, N., & M. Blackwell. 2001. Molecular phylogeny of dogwood anthracnose fungus (Discula destructiva) and the Diaporthales. Mycologia 93 (2): 355-365.

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