Morepork or boobook owl Ninox novaeseelandiae, from here.

Belongs within: Strigiformes.
Contains: Otus, Asio, Megascops, Bubo, Strix, Glaucidium.

The Striginae as recognised herein include all the living species of strigid owls (i.e. all living owls except barn owls and grass owls). Previous authors have divided the strigids between two sufamilies. The Striginae sensu stricto were characterised by a well-defined facial disc, and ear openings at least half the height of the skull, with a marginal dermal flap and a ligamentous or muscular bridge crossing the opening. The Buboninae, in contrast, lacked a strong facial disc and had smaller, simple ear openings (Kelso 1940). Recent phylogenies, however, have not supported this division, with the 'Striginae' being polyphyletic within the 'Buboninae'. Instead, the hawk-owls of the genus Ninox may diverge basally within the Strigidae; these relatively long-winged and -tailed owls are found in Asia and Australasia. Other genera include the boreal and saw-whet owls of the genus Aegolius, small, short-tailed and broad-winged owls with well-developed facial discs found in cooler regions of the Holarctic. Athene, the little owls, are also small, short-tailed species, but with less marked facial discs, found in Eurasia, north Africa and the Americas.

Striginae [Bubonidae, Buboninae]
    |--+--+--Otus JT12
    |  |  `--+--+--Asio JT12
    |  |     |  `--+--‘Otus’ leucotis JT12
    |  |     |     `--Ptilopsis granti JT12
    |  |     `--+--+--‘Otus’ flammeolus JT12
    |  |        |  `--Megascops JT12
    |  |        `--+--Bubo JT12
    |  |           `--+--Strix JT12
    |  |              `--+--Lophostrix cristata JT12
    |  |                 `--Pulsatrix JT12
    |  |                      |--P. koeniswaldiana JT12
    |  |                      |--P. melanota JT12
    |  |                      `--P. perspicillata JT12
    |  |                           |--P. p. perspicillata S55
    |  |                           `--P. p. saturata S55
    |  `--+--Micrathene whitneyi JT12
    |     `--+--Aegolius Kaup 1829 JT12, M02 [incl. Nyctale Brehm 1831 B94; Nyctalinae]
    |        |    |  i. s.: A. ridgwayi JT12
    |        |    |--A. funereus (Linnaeus 1758) JT12, M02 (see below for synonymy)
    |        |    |    |--A. f. funereus I92
    |        |    |    `--A. f. pallens (Schalow 1908) K92
    |        |    `--+--A. acadicus JT12
    |        |       `--A. harrisii JT12
    |        `--+--Glaucidium JT12
    |           `--Athene Boie 1822 JT12, M02 [incl. Noctura Savigny 1809 B94; Atheninae, Nocturinae]
    |                |  i. s.: A. angelis Mourer-Chauviré, Salotti et al. 1997 M02
    |                |         A. blewitti FP64
    |                |         A. cretensis Weesie 1982 M02
    |                |         A. murivora FP64
    |                |--A. cunicularia JT12
    |                `--+--A. brama JT12
    |                   `--A. noctua (Scopoli 1769) JT12, M02 (see below for synonymy)
    |                        |--A. n. noctua HR96
    |                        `--A. n. vidalii HR96
    `--Ninox Hodgson 1837 JT12, B94 [incl. Hieracoglaux Kaup 1848 B94, Ieraglaux Kaup 1851 B94; Ieraglaucinae]
         |  i. s.: N. affinis JT12
         |         N. burhani JT12
         |         N. ios JT12
         |         N. jacquinoti JT12
         |         N. meeki JT12
         |         N. ochracea JT12
         |         N. odiosa JT12
         |         N. punctulata JT12
         |         N. sumbaensis JT12
         |         N. superciliaris JT12
         |         N. theomacha JT12
         |         N. variegata JT12
         |--N. scutulata JT12
         |    |--N. s. scutulata M03
         |    `--N. s. japonica M03
         `--+--+--N. philippensis JT12
            |  `--+--N. rufa JT12
            |     |    |--N. r. rufa M03
            |     |    |--N. r. meesi M03
            |     |    `--N. r. queenslandica M03
            |     `--N. strenua JT12
            `--+--N. connivens (Latham 1801) JT12, WS48 [=Falco connivens WS48]
               |    |--N. c. connivens (see below for synonymy) WS48
               |    `--N. c. peninsularis M03
               `--+--N. natalis JT12 [=N. squamipila natalis M03]
                  `--+--N. rudolfi JT12
                     `--+--N. boobook JT12 [=N. novaeseelandiae boobook M03]
                        `--+--N. novaeseelandiae (Gmelin 1788) JT12, WS48 [=Strix novaeseelandiae WS48]
                           |    |--N. n. novaeseelandiae HWT01
                           |    |--N. n. albaria HR96
                           |    |--N. n. arida Mayr 1943 WS48
                           |    |--N. n. leucopsis M03
                           |    |--N. n. lurida M03
                           |    |--N. n. marmorata (Gould 1846) [=Athene marmorata] WS48
                           |    |--N. n. mixta Mathews 1912 [=N. boobook mixta] WS48
                           |    |--N. n. ocellata M03
                           |    `--N. n. undulata HWT01
                           `--N. squamipila M03

Striginae incertae sedis:
  Mioglaux Mlíkovský 1998 M02
    |--*M. debellatrix Mlíkovský 1998 M02
    `--M. poirrieri (Milne-Edwards 1863) [=Bubo poirrieri] M02
  Intulula Mlíkovský 1998 M02
    |--*I. tinnipara Mlíkovský 1998 M02
    `--I. brevis (Ballmann 1969) [=Strix brevis] M02
  Alasio Mlíkovský 1998 M02
    `--*A. collongensis (Ballmann 1972) [=Strix collongensis] M02
  Speotyto cunicularia RN72, A61
  Rhinoptynx clamator S18
  Ciccaba S18
    |--C. albogularis S55
    |--C. huhula S55
    |--C. nigrolineata S55
    |--C. virgata S55
    |    |--C. v. virgata FS55
    |    `--C. v. centralis FS55
    `--C. woodfordi S55
         |--C. w. woodfordi S55
         `--C. w. umbrina S55

Aegolius funereus (Linnaeus 1758) JT12, M02 [=Strix funerea M02; incl. Athene noctua veta Jánossy 1974 M02]

Athene noctua (Scopoli 1769) JT12, M02 [=Strix noctua M02; incl. A. noctua lunellensis Mourer-Chauviré 1975 M02]

Ninox connivens connivens (Latham 1801) [incl. N. connivens addenda Mathews 1912, N. connivens occidentalis Ramsay 1886] WS48

*Type species of generic name indicated


[A61] Austin, O. L., Jr. 1961. Birds of the World: A survey of the twenty-seven orders and one hundred and fifty-five families. Paul Hamlyn: London.

[B94] Bock, W. J. 1994. History and nomenclature of avian family-group names. Bulletin of the American Museum of Natural History 222: 1-281.

[FS55] Felten, H., & J. Steinbacher. 1955. Zur Vogelfauna von El Salvador. Senckenbergiana Biologica 36 (1-2): 9-19.

[FP64] Fisher, J., & R. T. Peterson. 1964. The World of Birds: A comprehensive guide to general ornithology. Macdonald: London.

[HR96] Heather, B. D., & H. A. Robertson. 1996. The Field Guide to the Birds of New Zealand. Viking: Auckland.

[HWT01] Holdaway, R. N., T. H. Worthy & A. J. D. Tennyson. 2001. A working list of breeding bird species of the New Zealand region at first human contact. New Zealand Journal of Zoology 28: 119-187.

[I92] Iwahashi, J. (ed.) 1992. Reddo Deeta Animaruzu: a pictorial of Japanese fauna facing extinction. JICC: Tokyo.

[JT12] Jetz, W., G. H. Thomas, J. B. Joy, K. Hartmann & A. Ø. Mooers. 2012. The global diversity of birds in space and time. Nature 491: 444-448.

Kelso, L. 1940. Variation of the external ear-opening in the Strigidae. Wilson Bulletin 52 (1): 24-29.

[M02] Mlíkovský, J. 2002. Cenozoic Birds of the World. Part 1: Europe. Ninox Press: Praha.

[M03] Morcombe, M. 2003. Field Guide to Australian Birds, 2nd ed. Steve Parish Publishing.

[RN72] Rutgers, A., & K. A. Norris (eds.) 1972. Encyclopaedia of Aviculture vol. 2. Blandford Press: London.

[S55] Steinbacher, J. 1955. Über die Schwanzmauser der Eulen (Strigidae) und Nachtschwalben (Caprimulgidae). Senckenbergiana Biologica 36: 235-240.

[S18] Stone, W. 1918. Birds of the Panama Canal Zone, with special reference to a collection made by Mr. Lindsey L. Jewel. Proceedings of the Academy of Natural Sciences of Philadelphia 70: 239-280.

[WS48] Whittell, H. M. & D. L. Serventy. 1948. A systematic list of the birds of Western Australia. Public Library, Museum and Art Gallery of Western Australia, Special Publication 1: 1-126.

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